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Cyrtodactylus sumuroi WELTON, SILER, LINKEM, DIESMOS & BROWN, 2010

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus sumuroi WELTON, SILER, LINKEM, DIESMOS & BROWN 2010 
DistributionPhilippines (Samar)

Type locality: Samar Island, Eastern Samar Province, Municipality of Taft, Barangay San Rafael, Taft Forest (11.829° N, 125.273° E; WGS84).  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: PNM 9723 (formerly KU 305566; field no. CDS 1834), adult male, collected on 16 June 2006 by CDS. 
DiagnosisDiagnosis: The critical comparisons for this species are congeners from the C. agusanensis complex. Cyrtodactylus sumuroi is distinguished from C. agusanensis by mid- body dorsals 87–98 (vs. 111–124); paraverte- brals 163–180 (vs. 184–196); postcloacal tubercles 4–6 (vs. 8–11); scales separating precloacals and femorals 19–21 (vs. 13–17); caudal annuli with tubercles 3–7 (vs. 7–12); canthal stripe present (vs. absent); anterior– posterior dorsal band projections minimal or absent (vs. extensive); and dark dorsal bands enclosing light bands (vs. light bands not enclosed). Cyrtodactylus sumuroi also exhibits tendencies toward fewer subdigital lamellae beneath Finger III (18–22 vs. 21–24), and midbody ventral scales (53–58 vs. 56–68).
Cyrtodactylus sumuroi is readily diagnosed from C. gubaot (Leyte) by SVL 73.9–84.9 (vs. 85.6–100.1), paravertebrals 163–180 (vs. 150– 162), ventral color cream (vs. dark gray with white flecks), anterior–posterior dorsal band projections minimal or absent (vs. moderate); dark dorsal bands enclosing light bands (vs. light bands not enclosed); cephalic tubercles moderate (vs. large); and number of caudal annuli with dorsal tubercles 3–7 (vs. 9–12). Cyrtodactylus sumuroi also exhibits tenden- cies toward fewer midbody ventrals (53–58 vs. 54–67), midbody dorsals (87–98 vs. 92–105), and subdigital lamellae beneath Finger III (18–22 vs. 20–26) and Toe IV (21–24 vs. 24– 30).
From C. mamanwa (Dinagat), C. sumuroi is distinguished by the presence (vs. absence) of a canthal stripe, anterior–posterior dorsal band projections minimal or absent (vs. moderate), cephalic tubercles moderate (vs. small), and caudal annuli with dorsal tubercles 3–7 (vs. 7–12). Cyrtodactylus sumuroi also exhibits tendencies toward fewer midbody dorsals (87–98 vs. 95–112), midbody ventrals (53–58 vs. 57–70), and paravertebrals (163– 180 vs. 173–192).
Cyrtodactylus sumuroi is diagnosable from all other Philippine congeners (C. annulatus, C. jambangan, C. tautbatorum, C. redimicu- lus, and C. philippinicus) by characters of body size and scalation (Table 3). The new species can be distinguished from C. jamban- gan and C. tautbatorum by larger SVL and the presence of dark dorsal bands enclosing light bands (vs. light bands not enclosed); from C. tautbatorum by a greater number of subdigital lamellae beneath Toe IV; from C. annulatus, C. jambangan, C. tautbatorum, and C. phi- lippinicus by the presence (vs. absence) of femoral pore-bearing scales; from C. tautba- torum and C. redimiculus by a greater number of midbody tubercle rows; from C. redimiculus by fewer paravertebral scales, fewer midbody dorsals, a greater number of dorsal tubercle rows, and larger cephalic tubercles; from C. annulatus, C. tautbatorum, C. redimiculus, and C. philippinicus by the presence of a canthal stripe (vs. absence); from C. annulatus by the presence of anterior–posterior dorsal band projections (vs. absence); and from C. annulatus, C. tautbatorum, and C. philippinicus by having fewer caudal annuli with dorsal tubercles (Table 3 in SILER et al. 2010). 
Comment 
EtymologyThe specific epithet is chosen in recognition of Juan Ponce (Agustin) Sumuroy, a Waray hero and popular Visayan leader of the ‘‘Sumuroy Rebellion’’ attempt of 1649. Although the Spanish suppressed the uprising and executed many of its leaders, by all accounts, Sumuroy and his coconspirators did much to spread the quest for independence among the peoples of the central Philippines during Spanish rule. 
References
  • Goldberg, S R & Bursey, C R; 2019. Cyrtodactylus sumuroi Endoparasites. Herpetological Review 50 (2): 372 - get paper here
  • Oaks, J. R., Siler, C. D. and Brown, R. M. 2019. The comparative biogeography of Philippine geckos challenges predictions from a paradigm of climate‐driven vicariant diversification across an island archipelago. Evolution doi:10.1111/evo.13754 - get paper here
  • Welton, Luke J.; Cameron D. Siler, Charles W. Linkem, Arvin C. Diesmos, and Rafe M. Brown 2010. Philippine Bent-Toed Geckos of the Cyrtodactylus agusanensis Complex: Multilocus Phylogeny, Morphological Diversity, and Descriptions of Three New Species. Herpetological Monographs 24 (1): 55-85 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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