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Cyrtodactylus bansocensis LUU, NGUYEN, LE, BONKOWSKI & ZIEGLER, 2016

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesLaotian: Ki Chiem Ban Soc
E: Ban Soc Bent-toed Gecko 
SynonymCyrtodactylus bansocensis LUU, NGUYEN, LE, BONKOWSKI & ZIEGLER 2016 
DistributionC Laos (Khammouane Province)

Type locality: on karst cliff, above the entrance of Peopalam cave (17°27.101’N, 105°35.393’E, 195 m elevation), near Ban Soc Village, Bualapha District, Khammouane Province, central Laos  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: VNUF (originally as VFU) R.2015.20, adult male, collected on 17 March 2015 by V. Q. Luu and K. Thanabuaosy. Paratype. NUOL R-2015.21, adult male, same collection data as the holotype. 
DiagnosisDiagnosis: As the original description of Cyrtodactylus bansocensis was based on two adult males (Luu et al. 2016c) we herein expand the diagnosis of this species as follows: medium size, SVL reaching 74.0 mm; dorsal pattern with four light transverse bands between limb insertions; supranasals in contact; 14–20 irregular dorsal tubercle rows at midbody; lateral skin fold present without tubercles; 34 or 35 ventral scale rows between ventrolateral folds; 34 precloacal and femoral pores in a continuous row in males (or interrupted by poreless scales); 16 precloacal and femoral pores in a continuous row in females (two outer femoral pores just pitted on each side), enlarged femoral and precloacal scales present; 4–7 postcloacal tubercles on each side; dorsal tubercles present at base of tail; subcaudal scales transversely enlarged (modified after Luu et al. 2016, Schneider et al. 2020). 
CommentSimilar species: C. rufford, C. khammouanensis, and C. soudthichaki. However, Cyrtodactylus bansocensis can be distinguished from C. rufford by having more ventral scale rows (34–35 versus 27–29), fewer supralabials (8–10 versus 10–12), fewer infralabials (8 versus 9–11), fewer femoral and precloacal pores in males (34 versus 42–43), and more postcloacal tubercles on each side (5–7 versus 4–5); C. khammouanensis by having fewer dorsal tubercle rows (14–15 versus 16–21), having fewer supralabials and infralabials (8–10 versus 11–12; 8 versus 9–10, respectively), fewer femoral and precloacal pores in males (34 versus 40–44), and tail with light rings (versus light bands); and from C. soudthichaki by its larger size (SVL reaching 74.0 mm versus 70.0 mm), having fewer dorsal tubercle rows (14–15 versus 19–20), more femoral and precloacal pores in males (34 versus 29), slightly higher number of ventral scales (34–35 versus 32–33), more subdigital lamellae on fourth toe (18–21 versus 18), more postcloacal tubercles on each side (5–7 versus 4–5), and tail with light rings (versus light bands) [Table 5 in LUU et al. 2016].

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyWe name this species after its type locality, Ban Soc limestone forest to underline the importance of this area (see also Ziegler et al. 2015) for biodiversity and nature conservation. 
References
  • Kwet, Axel 2017. Neue Arten: Liste der im Jahr 2016 neu beschriebenen Reptilien. Terraria-Elaphe 2017 (3): 54-70 - get paper here
  • LUU, VINH QUANG; Truong QUang Nguyen, MINH DUC LE, MICHAEL BONKOWSKI, THOMAS ZIEGLER 2016. A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa 4079 (1): 087–102 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • SCHNEIDER, NICOLE; VINH QUANG LUU, SALY SITTHIVONG, ALEXANDRE TEYNIÉ, MINH DUC LE, TRUONG QUANG NGUYEN, THOMAS ZIEGLER 2020. Two new species of Cyrtodactylus (Squamata: Gekkonidae) from northern Laos, including new finding and expanded diagnosis of C. bansocensis. Zootaxa 4822 (4): 503–530 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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