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Cyrtodactylus mombergi GRISMER, WOOD, QUAH, THURA, HERR & LIN, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Indawgyi Bent-toed Gecko 
SynonymCyrtodactylus mombergi GRISMER, WOOD, QUAH, THURA, HERR & LIN 2019 
DistributionMyanmar (Kachin: Indawgyi Wildlife Sanctuary)

Type locality: Yay Khan Kha stream east of Indawgyi Lake, Indawgyi Wildlife Sanctuary, Mohnyin Township, Kachin State, Myanmar (24.95694°N, 96.34690°E; 490 m in elevation).  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype. LSUHC 14734, Adult male, collected at 2100 h on 2 July 2018 by Evan S. H. Quah, Thura, M.K., Aung Ko Lin, Mark W. Herr, Perry L. Wood, Jr., Tun Tun Oo, Paw Lay, and L. Lee Grismer.
Paratypes. Adult females LSUHC 14730, 14732–33 and adult male LSUHC 14731 bear the same collection data as the holotype. Adult females LSUHC 14656–57 collected at 2230 h on 30 June 2018 by Thura, M.K., Aung Ko Lin, Evan S. H. Quah, Mark W. Herr, Perry L. Wood, Jr., Tun Tun Oo, Paw Lay, and L. Lee Grismer from Nant Tae stream east of Indawgyi Lake, Indawgyi Wildlife Sanctuary, Mohnyin Township, Kachin State, Myanmar (24.98732°N, 96.32352°E; 292 m in elevation). LSUHC 14698 collected during the evening by Mark W. Herr, Thura, M.K., Aung Ko Lin, Evan S. H. Quah, Perry L. Wood, Jr., Tun Tun Oo, Paw Lay, and L. Lee Grismer on 1 July 2018 from Wai Lone stream east of Indawgyi Lake, Indawgyi Wildlife Sanctuary, Mohnyin Township, Kachin State, Myanmar (24.94813°N, 96.33678°E; 499 m in elevation). 
DiagnosisDiagnosis: Cyrtodactylus mombergi sp. nov. differs from all other named species in the C. gansi group by having a maximum SVL of 74.0 mm; distinctly raised and keeled dorsal tubercles that extend beyond the base of the hemipenial swellings; 35–42 paravertebral tubercles; granular gular scales with no interspersed tubercles; a distinct ventrolateral body fold; 31–39 ventral scales; 19–22 fourth toe subdigital lamellae; no enlarged femoral scales or pores; no precloacal groove in males; 10 or 11 precloacal pores in males, 0–9 small pores in females; no dark band on nape; wide zig-zagged dorsal bands with paravertebral elements, and no lightened centers; dorsal bands not edged posteriorly with white tubercles; no white scales in ventrolateral fold; top of head usually darkly speckled to mottled; and 8–10 light-colored and 8–10 dark caudal bands. These characters are scored across all species of the C. gansi group (Table 2 in Grismer et al. 2019).


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CommentHabitat: Cyrtodactylus mombergi appears to be a nocturnal, scansorial, habitat generalist. All specimens were found in riverine evergreen forest in a wide variety of microhabitats including bamboo leaf-litter; rocky stream banks; on the vertical faces of granite boulders as high as 1.5 m above the ground; and on thin branches, tree limbs, and leaves of forest vegetation. 
EtymologyThis species is named in honor of Mr. Frank Momberg who was part of the initial field surveys of the Indawgyi region by FFI in January of 2010 that identified the watershed forests as important for the conservation of the Eastern Hoolock Gibbon (Hoolock leuconedys [Groves]). Mr. Momberg subsequently lead FFI’s efforts to conserve Indawgyi Lake and its watershed forests which resulted in the site being recognized as a Ramsar wetland of global importance in February of 2016 and a Man and Biosphere Reserve in June of 2017. Mr. Momberg continues to assist FFI in supporting the Man and Biosphere Reserve management board to assist local communities in addressing issues that threaten the natural values of the lake and surrounding forests in an effort to help them develop sustainable livelihoods. Additionally, Mr. Momberg has been unwavering in his support of our herpetological surveys throughout Myanmar. 
References
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., EVAN S. H. QUAH, MYINT KYAW THURA,<br>MARK W. HERR & AUNG KO LIN 2019. A new species of forest-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from the Indawgyi Wildlife Sanctuary, Kachin State, Myanmar. Zootaxa 4623 (1): 001–025 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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