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Cyrtodactylus muluensis DAVIS, BAUER, JACKMAN, NASHRIQ & DAS, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Mulu Bent-Toed Gecko 
SynonymCyrtodactylus muluensis DAVIS, BAUER, JACKMAN, NASHRIQ & DAS 2019 
DistributionEast Malaysia (Sarawak, Borneo)

Type locality: Lang Cave, Mulu National Park, Sarawak, East Malaysia. (04°1'34.34"N; 114°49'26.77"E; 155 m elevation; WGS 1984),  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: CAS 262983, adult male, collected by Izneil Nashriq and Hayden Davis on July 21, 2017 at 20:00–22:00 hrs.
Paratypes: CAS 262984; CAS 262986; (Figs. 1 & 2 in Davis et al. 2019)) have the same collection data as the holotype. Paratypes CAS 262995–262997 were collected from the karst surfaces situated between Clearwater Cave and Long Cave (04°3'47.048"N; 114°49'50.60"E; 205 m; WGS 1984), on July 23, 2017. 
DiagnosisDiagnosis: Cyrtodactylus muluensis sp. nov. can be differentiated from all other species of Cyrtodactylus by the combination of the following characters: maximum SVL of at least 88 mm; 10–13 supralabials; 8–11 infralabials; weak tuberculation on body; no tubercles on ventral surface of forelimbs, gular region, or ventrolateral folds; 33–54 paravertebral tubercles; 13–15 longitudinal tubercle rows; 33–42 ventral scales; 19–22 subdigital lamellae on fourth toe; 5 femoro-precloacal pores; enlarged median row of transverse scales; shallow precloacal groove in males; 5–8 dark dorsal body bands; body bands as wide as or slightly wider than interspaces; no rostral chevron; no white line edging the body bands and nuchal loop; banding pattern on dorsal side of body; no scattered white tubercles on dorsum; and nine dark caudal bands on original tail. These characters are scored across all currently described Bornean Cyrtodactylus species in Table 4 in Davis et al. 2019: 344.


Additional details (1542 characters) are available for collaborators and contributors. Please contact us for details. 
CommentSympatry: Cyrtodactylus consobrinus and C. cf. pubisulcus. However, C. consobrinus was seen almost exclusively on large tree trunks and C. cf. pubisulcus was almost exclusively on low-lying vegetation, indicating that they may not be syntopic with one another. 
EtymologyThe specific epithet muluensis is in reference to the type locality, Gunung Mulu National Park. As this is the first endemic gekkonid described from Gunung Mulu, the name muluensis is fitting. 
References
  • DAVIS, HAYDEN R.; AARON M. BAUER, TODD R. JACKMAN, IZNEIL NASHRIQ & INDRANEIL DAS 2019. Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia. Zootaxa 4614 (2): 331–352 - get paper here
  • Fukuyama R, Fukuyama I, Kurita T, Kojima Y, Hossman MY, Noda A, Nishikawa K 2021. New herpetofaunal records from Gunung Mulu National Park and its surrounding areas in Borneo. Herpetozoa 34: 89-96 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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