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Cyrtodactylus rivularis GRISMER, AOWPHOL, YODTHONG, AMPAI, TERMPRAYOON, AKSORNNEAM & RUJIRAWAN, 2022

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Pa La-U Bent-toed Gecko 
SynonymCyrtodactylus rivularis GRISMER, AOWPHOL, YODTHONG, AMPAI, TERMPRAYOON, AKSORNNEAM & RUJIRAWAN 2022: 130 
DistributionThailand (Prachuap Khiri Khan)

Type locality: Pa La-U Waterfall, Kaeng Krachan National Park, Huai Sat Yai Subdistrict, Hua Hin District, Prachuap Khiri Khan Province, Thailand (12.53685°N, 99.45972°E, 368 m a.s.l.).  
Reproduction 
TypesHolotype. Adult female ZMKU R 00947 collected by Attapol Rujirawan, Siriporn Yodthong, Korkhwan Termprayoon, Natee Ampai, and Piyawan Puanprapai on 15 September 2017.
Paratype. Adult female ZMKU R 00946 bearing the same data as the holotype. 
DiagnosisDiagnosis: Cyrtodactylus rivularis sp. nov. can be separated from all other species of the brevipalmatus group by the combination of having 12 or 13 supralabials, 9–11 infralabials, 33 or 34 paravertebral tubercles, 18–20 rows of longitudinally arranged tubercles, 34–37 transverse rows of ventrals, 160–166 longitudinal rows of ventrals, nine expanded subdigital lamellae on the fourth toe, 12 or 13 unexpanded subdigital lamellae on the fourth toe, 21 or 22 total subdigital lamellae on the fourth toe; eight expanded subdigital lamellae on the fourth finger, 10–12 unexpanded subdigital lamellae on the fourth finger, 18–20 total subdigital lamellae on the fourth finger; 14–16 total enlarged femoral scales; 15 enlarged precloacals; enlarged femorals and enlarged precloacals not continuous, and lacking pores; proximal femorals less than one-half the size of the distal femorals; small tubercles on forelimbs and flanks; large dorsolateral caudal tubercles and wide ventrolateral caudal fringe; ventrolateral caudal fringe composed generally homogeneous scales; tail square in cross-section; single enlarged unpaired medial subcaudals not posteromedially furrowed; maximum SVL 73.9 mm; three or four dark transverse body bands (Tables 4, 5). (Grismer et al. 2022)


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Comment 
EtymologyThe specific epithet rivularis is derived from the Latin rivus, meaning stream, brook, or creek refers to rocky brook or stream habitat of the new species. 
References
  • Grismer LL, Aowphol A, Yodthong S, Ampai N, Termprayoon K, Aksornneam A, Rujirawan A 2022. Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162 - get paper here
  • Grismer LL, Rujirawan A, Chomdej S, Suwannapoom C, Yodthong S, Aksornneam A, Aowphol A 2023. A new species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) from the uplands of western Thailand. ZooKeys 1141: 93–118 - get paper here
  • GRISMER, L. LEE; PARINYA PAWANGKHANANT, ANDREY M. BRAGIN, ALEXEY V. TROFIMETS, ROMAN A. NAZAROV, CHATMONGKON SUWANNAPOOM & NIKOLAY A. POYARKOV 2024. A new species of the Cyrtodactylus brevipalmatus group (Squamata: Gekkonidae) from Phuket Island, Thailand with a discussion of the group’s potential biogeography. Zootaxa 5437 (2): 193–222 - get paper here
  • Kwet, A. 2023. Liste der im Jahr 2022 neu beschriebenen Reptilien. Elaphe 2023 (3): 48-73
 
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