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Cyrtodactylus sanpelensis GRISMER, WOOD, THURA, ZIN, QUAH, MURDOCH, GRISMER, LIN, KYAW & LWIN, 2017

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Sanpel cave bent-toed gecko 
SynonymCyrtodactylus sanpelensis GRISMER, WOOD, THURA, ZIN, QUAH, MURDOCH, GRISMER, LIN, KYAW & LWIN 2017: 58 
DistributionMyanmar (Mawlamyine District, Mon State)

Type locality: Sanpel Cave 21.3 km south-east of Mawlamyine, Mawlamyine District, Mon State, Myanmar (N16°22.427, E97°46.388; 44 m in elevation).  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 12877, Adult male collected on 8 October 2016 between 1600 and 2000 h by Myint Kyaw Thura, L. Lee Grismer, Marta S. Grismer, Evan S. H. Quah, Perry L. Wood, Jr., Matthew L. Murdoch, Thaw Zin and Htet Kyaw. Paratypes: Adult males BYU 52223, LSUHC 12875, 12881, 12883, 12887, 12889–90, adult females BYU 52221–22, 52224, LSUHC 12878–80, 12886 and juvenile LSUHC 12885 bear the same collection data as the holotype. 
DiagnosisDiagnosis: Cyrtodactylus sanpelensis sp. nov. differs from all congeners by having the unique combination of 9–11 supralabials; seven or eight infralabials; 12–14 rows of longitudinal body tubercles; 27–31 paravertebral tubercles; 35–41 ventral scales; relatively long digits with eight or nine expanded subdigital lamellae proximal to the digital inflection on the fourth toe, 12 or 13 unmodified distal subdigital lamellae, 20–22 total subdigital lamellae; low, weakly keeled, dorsal body tubercles; body tubercles not extending beyond base of tail; enlarged femoral and precloacal scales not continuous; 14–20 enlarged femoral scales; 8–16 femoral pores in males; 9–12 enlarged precloacal scales; seven or eight precloacal pores in males; two rows of enlarged post-precloacal scales; median subcaudal scales three times as wide as long, extending onto lateral surface of tail; top of head diffusely mottled, no yellow reticulum; nuchal loop not divided medially, lacking an anterior, azygous notch, posterior border straight or sinuous; five (rarely four), dark, regularly shaped, dorsal bands not wider than interspaces, nearly always with lightened centres, not edged with white tubercles; band on nape present; dark markings in dorsal interspaces; ventrolateral fold not whitish; anterodorsal margin of thighs and brachia unpigmented; 7–10 light caudal bands bearing dark markings, not encircling tail; 7–10 dark caudal bands wider than light caudal bands; and regenerated tail not spotted.


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Comment 
EtymologyThe specific epithet, sanpelensis, is a noun in apposition in reference to the type locality of Sanpel Cave. 
References
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR., MYINT KYAW THURA, THAW ZIN, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MARTA S. GRISMER, AUNG LIN, HTET KYAW and NGWE LWIN 2017. Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in east- central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism. Zoological Journal of the Linnean Society 182: 862-959 - get paper here
  • Grismer, Lee; Wood, Perry L., Jr.; Poyarkov, Nikolay A.; Le, Minh D.; Karunarathna, Suranjan; Chomdej, Siriwadee; Suwannapoom, Chatmongkon; Qi, Shuo; Liu, Shuo; Che, Jing; Quah, Evan S.H.; Kraus, Fred; Oliver, Paul M.; Riyanto, Awal; Pauwels, Olivier 2021. Karstic Landscapes Are Foci of Species Diversity in the World’s Third-Largest Vertebrate Genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata; Gekkonidae). Diversity 13 (5): 183 - get paper here
  • Riedel, J., Eisele, K., Gabelaia, M., Higham, T. E., Wu, J., Do, Q. H., Nguyen, T. Q., Meneses, C. G., Brown, R. M., Ziegler, T., Grismer, L. L., Russell, A. P., & Rödder, D. 2024. Ecologically-related variation of digit morphology in Cyrtodactylus (Gekkota, Squamata) reveals repeated origins of incipient adhesive toepads. Functional Ecology, 00, 1–19 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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