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Cyrtodactylus muluensis DAVIS, BAUER, JACKMAN, NASHRIQ & DAS, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Mulu Bent-Toed Gecko 
SynonymCyrtodactylus muluensis DAVIS, BAUER, JACKMAN, NASHRIQ & DAS 2019 
DistributionEast Malaysia (Sarawak, Borneo)

Type locality: Lang Cave, Mulu National Park, Sarawak, East Malaysia. (04°1'34.34"N; 114°49'26.77"E; 155 m elevation; WGS 1984),  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: CAS 262983, adult male, collected by Izneil Nashriq and Hayden Davis on July 21, 2017 at 20:00–22:00 hrs.
Paratypes: CAS 262984; CAS 262986; (Figs. 1 & 2 in Davis et al. 2019)) have the same collection data as the holotype. Paratypes CAS 262995–262997 were collected from the karst surfaces situated between Clearwater Cave and Long Cave (04°3'47.048"N; 114°49'50.60"E; 205 m; WGS 1984), on July 23, 2017. 
DiagnosisDiagnosis: Cyrtodactylus muluensis sp. nov. can be differentiated from all other species of Cyrtodactylus by the combination of the following characters: maximum SVL of at least 88 mm; 10–13 supralabials; 8–11 infralabials; weak tuberculation on body; no tubercles on ventral surface of forelimbs, gular region, or ventrolateral folds; 33–54 paravertebral tubercles; 13–15 longitudinal tubercle rows; 33–42 ventral scales; 19–22 subdigital lamellae on fourth toe; 5 femoro-precloacal pores; enlarged median row of transverse scales; shallow precloacal groove in males; 5–8 dark dorsal body bands; body bands as wide as or slightly wider than interspaces; no rostral chevron; no white line edging the body bands and nuchal loop; banding pattern on dorsal side of body; no scattered white tubercles on dorsum; and nine dark caudal bands on original tail. These characters are scored across all currently described Bornean Cyrtodactylus species in Table 4 in Davis et al. 2019: 344.


Additional details (1542 characters) are available for collaborators and contributors. Please contact us for details. 
CommentSympatry: Cyrtodactylus consobrinus and C. cf. pubisulcus. However, C. consobrinus was seen almost exclusively on large tree trunks and C. cf. pubisulcus was almost exclusively on low-lying vegetation, indicating that they may not be syntopic with one another. 
References
  • DAVIS, HAYDEN R.; AARON M. BAUER, TODD R. JACKMAN, IZNEIL NASHRIQ & INDRANEIL DAS 2019. Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia. Zootaxa 4614 (2): 331–352 - get paper here
  • Fukuyama R, Fukuyama I, Kurita T, Kojima Y, Hossman MY, Noda A, Nishikawa K 2021. New herpetofaunal records from Gunung Mulu National Park and its surrounding areas in Borneo. Herpetozoa 34: 89-96 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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