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Cyrtodactylus chaunghanakwaensis GRISMER, WOOD, THURA, QUAH, MURDOCH, GRISMER, HERR, LIN & KYAW, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Chaunghanakwa Hill bent-toed gecko 
SynonymCyrtodactylus chaunghanakwaensis GRISMER, WOOD, THURA, QUAH, MURDOCH, GRISMER, HERR, LIN & KYAW 2018 
DistributionMyanmar (Mon: Mawlamyine District)

Type locality: Chaunghanakwa Hill, 50 km south-east of Mawlamyine, Mawlamyine District, Mon State, Myanmar (16.18456°N, 97.98814°E; 14 m in elevation  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 13305, Adult male, collected on 13 May 2017 at 2000 hrs by Myint Kyaw Thura, Mark W. Herr, L. Lee Grismer, Perry L. Wood, Jr., Matthew L. Murdoch, Evan S.H. Quah, and Htet Kyaw.
Paratypes: Adult male paratypes LSUHC 13295, 13298, 13300, 13302, 13308, and 13312–14 and adult female paratypes LSUHC 13294, 13296–97, 13299, 13301, 13303–04, 13306–07, 13309–11, 13315–16, 13321 bear the same collecting data as the type series. 
DiagnosisDiagnosis: Cyrtodactylus chaunghanakwaensis sp. nov. differs from all congeners by having the unique combination of a maximum SVL of 76.3 mm; 8–11 supralabials; 7–9 infralabials; 31–36 paravertebral tubercles; 17–22 longitudinal rows of body tubercles; 23–27 long- itudinal rows of ventral scales ventral scales; relatively long digits with 6–9 expanded subdigital lamellae proximal to the digital inflexion on the fourth toe, 11–14 unmodified distal subdigital lamellae, 18–21 total subdigital lamellae; dorsal body tubercles low, weakly keeled in adults; tubercles extendind beyond base of tail; enlarged femoral and precloacal scales continuous; enlarged proximal femoral scales greater than one-half the size of enlarged distal femoral scales; 27–32 enlarged femoral scales; 27–32 femoral pores in males; 9–11 precloacal pores in males; femoral and precloacal pore-bearing scales continuous; 9–12 enlarged precloacal scales; two or three rows of enlarged post- precloacal scales; medial subcaudal scales 3 times as wide as long, extending onto lateral surface of tail; top of head darkly mottled, no yellow reticulum; nuchal loop not divided medially, no pronounced anterior azygous notch, posterior border jagged; 5–7 jagged, dark, dorsal bands bearing paravertebral elements, wider than interspaces with variably lightened centres, variably edged with white tubercles; nape band present; dark mark- ings in dorsal interspaces; ventrolateral folds not whitish; anterodrosal margins of thighs and brachia darkly pigmented; 10–14 light caudal bands bearing dark markings, not encircling tail; 11–15 dark caudal bands wider than light caudal bands; and mature regenerated tail not spotted. These characters are scored against all other species in the sinyineensis group (Table 4) and all other Burmese species of the Indochinese clade in Grismer et al. (2017a, table 8).


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Comment 
EtymologyThe specific epithet, chaunghanakwaensis, is a noun in apposition in reference to the type locality of Chaunghahakwa Hill. 
References
  • CHOMDEJ, S., SUWANNAPOOM, C., PAWANGKHANANT, P., PRADIT, W., NAZAROV, R. A., GRISMER, L. L., & POYARKOV, N. A. 2020. A new species Cyrtodactytlus Gray (Squamata: Gekkonidae) from western Thailand and the phylogenetic placement of C. inthanon and C. doisuthep. Zootaxa 4838 (2): 179-209 - get paper here
  • Grismer, L. Lee; Perry L. Wood Jr., Myint Kyaw Thura, Evan S.H. Quah, Matthew L. Murdoch, Marta S. Grismer, Mark W. Herr, Aung Lin & Htet Kyaw 2018. Three more new species of Cyrtodactylus (Squamata: Gekkonidae) from the Salween Basin of eastern Myanmar underscore the urgent need for the conservation of karst habitats. Journal of Natural History, 52:19-20, 1243-1294 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., EVAN S.H. QUAH, MARTA S. GRISMER, MYINT KYAW<br>THURA, JAMIE R. OAKS & AUNG LIN 2020. Two new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from a karstic archipelago in the Salween Basin of southern Myanmar (Burma). Zootaxa 4718 (2): 151–183 - get paper here
  • Keating, S. E., Blumer, M., Grismer, L. L., Lin, A., Nielsen, S. V., Thura, M. K., ... & Gamble, T. 2021. Sex Chromosome Turnover in Bent-Toed Geckos (Cyrtodactylus). Genes, 12(1), 116 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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