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Cyrtodactylus markuscombaii (DAREVSKY, HELFENBERGER, ORLOV & SHAH, 1998)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Striped gecko 
SynonymGonydactylus markuscombaii DAREVSKY, HELFENBERGER, ORLOV & SHAH 1998: 91
Gymnodactylus marcuscombaii
Cyrtopodion markuscombaii — RÖSLER 2000: 74
Siwaligekko markuscombaii — KHAN 2003
Cyrtodactylus marcuscombaii — MAHONY et al. 2009 
DistributionE Nepal (Mechi), (only known from type locality)

Type locality: road between Ilam town and Pawakhola village, 1200-1300 m elevation), Ilam distr. Mecha Prov., E Nepal (Ilam: 26°54'54''N, 87°55'48''E; Pawakhola: 26°58'33''N, 87°54'39''E).  
Reproductionoviparous 
TypesHolotype: MHNG 2590.35, adult male; paratype: ZISP 20685 
DiagnosisDiagnosis: A medium-sized species of Gonydactylus with an adult body size up to 70-72 mm in SVL; no preanal groove; femoral pores and enlarged femoral scales absent; preanal pores present; dorsal surface of the body with fine granules arranged into irregular longitudinal rows; scale rows between the distinct ventrolateral folds 38-39; tail not segmented; transversely widened subcaudal scales ab sent; dark irregular spots on the dorsum present; tail longer than snout-vent length, the end-limbs not reaching the axilla (Darevsky et al. 1997). 
CommentSynonymy: Gonydactylus KUHL & VAN HASSELT 1822 is a nomen nudum.

Authorship: note that one of the authors was misspelled on the original paper as “Helfeberger” instead of Helfenberger. 
EtymologyNamed after Markus Comba (b.1956), a Swiss antiquarian book-seller and book-restorer, who is also a naturalist. He supported several expeditions to Central Asia and to Nepal. 
References
  • AGARWAL, ISHAN; STEPHEN MAHONY, VARAD B. GIRI, R. CHAITANYA, AARON M. BAUER 2018. Six new Cyrtodactylus (Squamata: Gekkonidae) from northeast India. Zootaxa 4524 (5): 501–535 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bhattarai S., Gurung A., Lamichhane BR., Regmi R., Dhungana M., Kumpakha B. and Subedi N. 2020. Amphibians and Reptiles of Chure Range, Nepal. President Chure Terai-Madhesh Conservation Development Board and National Trust for Nature Conservation, Khumaltar, Lalitpur, Nepal
  • Darevsky, I. S., N. Helfenberger*, N. Orlov & K. Shah 1998. Two new species of the genus Gonydactylus (Sauria: Gekkonidae) from eastern Nepal. Russ. J. Herpetol. 4 (2): 89-93 [1997; * misspelled as “Helfeberger” on the original paper] - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Kästle , W., Rai, K. & Schleich, H.H. 2013. FIELD GUIDE to Amphibians and Reptiles of Nepal. ARCO-Nepal e.V., 625 pp. - get paper here
  • Khan, M.S. 2003. Questions of generic designation of angular-toed geckos of Pakistan with descriptions of three new genera (Reptilia: Gekkonidae). J. nat. hist. Wildl. 2 (2): 1-9
  • Mahony, S., M. Ahmed, M. K. Hossain, M. M. Kabir & M. K. Hasan 2009. Cyrtodactylus ayeyarwadyensis Bauer, 2003 (Reptilia: Gekkonidae) in Bangladesh with habitat details of new collection localities and a discussion of morphological variation. Salamandra 45 (4): 245–260 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
 
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