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Cyrtodactylus kulenensis GRISMER, GEISSLER, NEANG, HARTMANN, WAGNER & POYARKOV, 2021

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus kulenensis GRISMER, GEISSLER, NEANG, HARTMANN, WAGNER & POYARKOV 2021
Cyrtodactylus cf. intermedius — GEISSLER et al. 2019 
DistributionCambodia (Siem Reap Province, Banteay Srei District)

Type locality: Phnom Kulen National Park, Phnom Kbal, Banteay Srei District, Siem Reap Province, Cambodia (13°41’21’’N, 104°00’55’’E°N at about 490 m in elevation)  
Reproduction 
TypesHolotype: ZFMK 92573, adult male, collected by T. Hartmann and P. Geissler on 2 June 2011
Paratypes: ZFMK 88356, ZFMK 88357, ZFMK 88359, adult females and subadult male ZFMK 88358, collected by T. Hartmann in September 2008. CBC 03030, adult female and ZFMK 90311, juvenile, collected by T. Hartmann in June 2009. ZFMK 92572, ZFMK 92574, ZFMK 92575, adult females, collected by T. Hartmann and P. Geissler in June 2011. 
DiagnosisDiagnosis: Cyrtodactylus kulenensis can be separated from all other species of the C. intermedius group by having 8–11 supralabials, 8–11 infralabials, 33–38 paravertebral tubercles, 17–19 rows of longitudinally arranged tubercles, 38–44 ventrals, seven or eight expanded subdigital lamellae on the fourth toe, 11–13 unexpanded subdigital lamellae on the fourth toe, 18–21 total subdigital lamellae on the fourth toe, 12–21 enlarged femorals, nine or 10 enlarged precloacals, two or three rows of enlarged post-precloacals, two or three postcloacal tubercles, nine or 10 precloacal pores in males (N=2), enlarged femorals and enlarged precloacals continuous, proximal femorals less than one-half the size of the distal femorals, no digital pockets, maximum SVL 89.6 mm, no large dark blotches on top of head, three or four well-defined brown body bands edged with white tubercles, and body bands narrower than the beige interspaces (Grismer et al. 2021).


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Comment 
EtymologyThe specific epithet kulenensis is a Latinized toponymic adjective named after Phnom (=Mount) Kulen National Park, Banteay Srei District, Siem Reap Province, Cambodia. “Phnom Kulen”means “the Mountain of Lychees” in Khmer language. 
References
  • CHHIN, S., NEANG, T., CHAN, S., KONG, K., OU, R., SAMORN, V., ... & GRISMER, L. L. 2024. A new species in the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from an isolated limestone karst formation in southwestern Cambodia. Zootaxa 5474 (1): 1-20
  • GRISMER, L. LEE; PETER GEISSLER, THY NEANG, TIMO HARTMANN, PHILIPP WAGNER, & NIKOLAY A. POYARKOV 2021. Molecular phylogenetics, PCA, and MFA recover a new species of Cyrtodactylus (Squamata: Gekkonidae) from an isolated sandstone massif in northwestern Cambodia. Zootaxa 4949 (2): 261–288 - get paper here
  • Grismer, Lee; Wood, Perry L., Jr.; Poyarkov, Nikolay A.; Le, Minh D.; Karunarathna, Suranjan; Chomdej, Siriwadee; Suwannapoom, Chatmongkon; Qi, Shuo; Liu, Shuo; Che, Jing; Quah, Evan S.H.; Kraus, Fred; Oliver, Paul M.; Riyanto, Awal; Pauwels, Olivier 2021. Karstic Landscapes Are Foci of Species Diversity in the World’s Third-Largest Vertebrate Genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata; Gekkonidae). Diversity 13 (5): 183 - get paper here
  • Riedel, J., Eisele, K., Gabelaia, M., Higham, T. E., Wu, J., Do, Q. H., Nguyen, T. Q., Meneses, C. G., Brown, R. M., Ziegler, T., Grismer, L. L., Russell, A. P., & Rödder, D. 2024. Ecologically-related variation of digit morphology in Cyrtodactylus (Gekkota, Squamata) reveals repeated origins of incipient adhesive toepads. Functional Ecology, 00, 1–19 - get paper here
 
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