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Cyrtodactylus martinstolli (DAREVSKY, HELFENBERGER, ORLOV & SHAH, 1998)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Ilam bent-toed gecko, Barred gecko
E: Striped gecko [markuscombaii] 
SynonymGonydactylus martinstolli DAREVSKY, HELFENBERGER, ORLOV & SHAH 1998: 89
Gonydactylus markuscombaii DAREVSKY, HELFENBERGER, ORLOV & SHAH 1998: 91
Cyrtodactylus martinstollii — DAREVSKY et al. 1997
Gymnodactylus martinstollii — DAREVSKY et al. 1997
Cyrtopodion martinstolli — RÖSLER 2000: 74
Cyrtopodion markuscombaii — RÖSLER 2000: 74
Siwaligekko markuscombaii — KHAN 2003
Siwaligekko martinstolli — KHAN 2003
Cyrtodactylus martinstollii — MAHONY et al. 2009
Cyrtodactylus marcuscombaii — MAHONY et al. 2009
Cyrtodactylus martinstollii — BHATTARAI et al. 2025: 409 
DistributionNepal (only known from type locality)

Type locality: road between Ilam town and Pawakhola village, 1200-1300 m elevation, Ilam distr. Mecha Prov., E Nepal (Ilam: 26°54'54''N, 87°55'48''E; Pawakhola: 26°58'33''N, 87°54'39''E)

markuscombaii (invalid): E Nepal (Mechi), (only known from type locality); Type locality: road between Ilam town and Pawakhola village, 1200-1300 m elevation), Ilam distr. Mecha Prov., E Nepal (Ilam: 26°54'54''N, 87°55'48''E; Pawakhola: 26°58'33''N, 87°54'39''E).  
Reproductionoviparous 
TypesHolotype: MHNG 2590.09; paratypes: MHNG (23): 2590.10 – 13; 2590.14 – 20; 2590.21 – 33; ZISP 20685
Holotype: MHNG 2590.35, adult male; paratype: ZISP 20685 [markuscombaii] 
DiagnosisDiagnosis (Tables 2 & 3). A medium-sized Cyrtodactylus, snout to vent length up to 80.4 mm (n=7). Dorsal pholidosis heterogeneous; smooth granular scales intermixed with fairly regularly arranged rows of enlarged, feebly keeled to smooth, weakly pointed tubercles; a ventrolateral fold of skin present on lower flank; 19–23 rows of dorsal tubercles at midbody, 30–37 tubercles in paravertebral rows; ventral scales subequal from chest to vent, smooth, subcircular, and subimbricate with rounded end; 35–40 scales across belly at midbody, 84–91 longitudinal scales between axilla to groin, 184–196 longitudinal scales from mental to cloaca; subdigital scansors smooth, unnotched, and mostly entire; 14 or 15 lamellae under digit I of manus and pes, 18–20 lamellae under digit IV of manus and 19–22 lamellae under digit IV of pes; males with continuous series of seven or eight precloacal pores and 9–12 enlarged precloacal scales on pore bearing scale row (n=3); female without pores but having eight or nine pitted scales, and 8–10 enlarged precloacal scales on pit bearing scale row (n=4); scales on non-regenerated tail dorsum homogeneous; composed of fairly regularly arranged, smooth, subcircular, flattened, and subimbricate scales that are larger than granular scales on midbody dorsum, gradually becoming larger posteriorly and dorsolaterally; a few scattered enlarged tubercles present on the tail base; subcaudal scales in median series smooth, variable in size and shape, and not enlarged; variegated dorsal pattern with 8–11 dark bands, original tail with 10–12 alternating dark and lighter bands. (Bhattarai et al. 2025)


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CommentAuthorship: note that one of the authors was misspelled on the original paper as “Helfeberger” instead of Helfenberger.

Synonymy: Gonydactylus KUHL & VAN HASSELT 1822 is a nomen nudum. Bhattarai et al. 2025 synonymized C. markuscombai with C. martinstolli.

RDB speciesID: 13051 [markuscombaii]

Etymology: C. markuscombaii was named after Markus Comba (b.1956), a Swiss antiquarian book-seller and book-restorer, who is also a naturalist. He supported several expeditions to Central Asia and to Nepal. 
References
  • AGARWAL, ISHAN; STEPHEN MAHONY, VARAD B. GIRI, R. CHAITANYA, AARON M. BAUER 2018. Six new Cyrtodactylus (Squamata: Gekkonidae) from northeast India. Zootaxa 4524 (5): 501–535 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bhattarai S., Gurung A., Lamichhane BR., Regmi R., Dhungana M., Kumpakha B. and Subedi N. 2020. Amphibians and Reptiles of Chure Range, Nepal. President Chure Terai-Madhesh Conservation Development Board and National Trust for Nature Conservation, Khumaltar, Lalitpur, Nepal
  • BHATTARAI, S., GAUTAM, B., NEUPANE, B. P., KHANDEKAR, A., THACKERAY, T., AGARWAL, I., ... & WRIGHT, W. 2025. A review of the genus Cyrtodactylus Gray 1827 (Squamata: Gekkonidae) of Nepal with descriptions of three new species. Zootaxa 5594(3): 401-451 - get paper here
  • Darevsky, I. S., N. Helfenberger*, N. Orlov & K. Shah 1998. Two new species of the genus Gonydactylus (Sauria: Gekkonidae) from eastern Nepal. Russ. J. Herpetol. 4 (2): 89-93 [1997; * misspelled as “Helfeberger” on the original paper] - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Kästle , W., Rai, K. & Schleich, H.H. 2013. FIELD GUIDE to Amphibians and Reptiles of Nepal. ARCO-Nepal e.V., 625 pp. - get paper here
  • Khan, M.S. 2003. Questions of generic designation of angular-toed geckos of Pakistan with descriptions of three new genera (Reptilia: Gekkonidae). J. nat. hist. Wildl. 2 (2): 1-9
  • Mahony, S., M. Ahmed, M. K. Hossain, M. M. Kabir & M. K. Hasan 2009. Cyrtodactylus ayeyarwadyensis Bauer, 2003 (Reptilia: Gekkonidae) in Bangladesh with habitat details of new collection localities and a discussion of morphological variation. Salamandra 45 (4): 245–260 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Uetz, Peter; Tapil Prakash Rai, Ruma Thapa, Barbara Lester & Paul Freed 2022. Herpetologische Reise nach Nepal. [A Herpetological trip to Nepal] Sauria 44 (4): 3-31 - get paper here
 
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