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Cyrtodactylus pseudoquadrivirgatus RÖSLER, NGUYEN, VU, NGO & ZIEGLER, 2008

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus pseudoquadrivirgatus RÖSLER, NGUYEN, VU, NGO & ZIEGLER 2008
Cyrtodactylus cf. pseudoquadrivirgatus — JESTRZEMSKI et al. 2013
Cyrtodactylus pseudoquadrivirgatus — NGUYEN et al. 2021 
DistributionVietnam (Thua Thien-Hue Province)

Type locality: A Luoi, Thua Thien-Hue Province, central Vietnam, at an elevation of 800 m above sea level.  
Reproductionoviparous 
TypesHolotype: ZFMK 83895, adult female, (Fig. 1–2a– d), leg. Vu Ngoc Thanh, 20 July 2005. 
DiagnosisDiagnosis. A medium-sized Cyrtodactylus that can be distinguished from all congeners from the Oriental and Australian regions on the basis of the following combination of characters: 1) SVL up to 84 mm (TL > SVL); 2) head depressed, distinct from body, snout with me dial concavity; 3) 37–55 interorbital scales; 4) body rounded in cross-section, belly lat, not wider than head; 5) lateral fold narrow, with enlarged lateral tubercles; 6) tubercles present on dorsum of head, body, forearm, hind limbs and tail; 7) 16–24 rows of dorsal tubercles; 8) 41–57 longitudinal rows of ventral scales at midbody; 9) rounded tail in cross-section, tail base not enlarged; 10) 5–9 precloacal pores in angular series and 1–12 enlarged precloacal scales; 11) absence of enlarged femoral scales, lack of femoral pores and a precloacal groove; 12) absence of transversely enlarged subcaudals; 13) 9–13 lamellae under first digit, 10–13 lamellae under irst toe, and 16–25 lamellae under 4th toe; 14) body dorsum light brown with dark brown pattern (head mottled, dorsum mottled, striped or with bands); 15) neck band medially interrupted; 16) limbs striped or mottled, tail with dark and light bands (Rösler et al. 2008).


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Comment 
EtymologyThe name refers to Cyrtodactylus quadrivirgatus, with which the new species may be easily confused with due to the striking similarities in external morphology, including scalation, colouration and pattern; the prefix pseudo derives from the Greek pseudes = false, pretended. 
References
  • DAVID, PATRICK; TRUONG QUANG NGUYEN, NICOLE SCHNEIDER & THOMAS ZIEGLER 2011. A new species of the genus Cyrtodactylus Gray, 1827 from central Laos (Squamata: Gekkonidae). Zootaxa 2833: 29–40 - get paper here
  • Geissler, P.; Nazarov, R.; Orlov, N.L.; Böhme, W.; My Phung, T.; Nguyen, T.Q. & Ziegler, T. 2009. A new species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa 2161: 20–32 - get paper here
  • Jestrzemski, Daniel; Stefan Schütz, Truong Quang Nguyen, Thomas Ziegler 2013. A survey of amphibians and reptiles in Chu Mom Ray National Park, Vietnam, with implications for herpetofaunal conservation. Asian Journal of Conservation Biology, 2 (2): 88–110 - get paper here
  • LUU, VINH QUANG; THOMAS CALAME, Truong QUang Nguyen, MICHAEL BONKOWSKI & THOMAS ZIEGLER 2015. A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa 4058 (3): 388–402 - get paper here
  • LUU, VINH QUANG; TRUONG QUANG NGUYEN, HUY QUANG DO, THOMAS ZIEGLER, 2011. A new Cyrtodactylus (Squamata: Gekkonidae) from Huong Son limestone forest, Hanoi, northern Vietnam. Zootaxa 3129: 39–50 - get paper here
  • LUU, VINH QUANG; Truong QUang Nguyen, MINH DUC LE, MICHAEL BONKOWSKI, THOMAS ZIEGLER 2016. A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa 4079 (1): 087–102 - get paper here
  • LUU, VINH QUANG;THOMAS CALAME, MICHAEL BONKOWSKI, TRUONG QUANG NGUYEN & THOMAS ZIEGLER 2014. A new species of Cyrtodactylus (Squamata: Gekkonidae) from Khammouane Province, Laos. Zootaxa 3760 (1): 054–066 - get paper here
  • NAZAROV, ROMAN; NIKOLAY A. POYARKOV, NIKOLAI L. ORLOV, TRUNG MY PHUNG, TAO THIEN NGUYEN, DUC MINH HOANG & THOMAS ZIEGLER 2012. Two new cryptic species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa 3302: 1–24 - get paper here
  • Ngo van Tri, L. Lee Grismer & J. L. Grismer 2010. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Phu Quoc National Park, Kien Giang Biosphere Reserve, Southwestern Vietnam. Zootaxa 2604: 37–51 - get paper here
  • Ngo, Van Tri 2011. Cyrtodactylus martini, another new karst-dwelling Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Northwestern Vietnam. Zootaxa 2834: 33–46 - get paper here
  • NGO, VAN TRI & L. LEE GRISMER 2012. A new endemic species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Tho Chu Island, southwestern Vietnam. Zootaxa 3228: 48–60 - get paper here
  • Nguyen AT, Duong TV, Grismer LL, Poyarkov NA 2021. A new granite cave-dwelling Bent-toed Gecko from Vietnam of the Cyrtodactylus irregularis group (Squamata; Gekkonidae) and a discussion on cave ecomorphology. Vertebrate Zoology, 71: 155-174 - get paper here
  • Nguyen, Luan Thanh; Ha Van Hoang, Thang Tai Nguyen, McCormack Timothy E.M., Sang Ngoc Nguyen 2016. A COLLECTION OF AMPHIBIANS AND REPTILES FROM BAC HUONG HOA NATURE RESERVE, QUANG TRI PROVINCE, VIETNAM. Proceedings of the 3nd National Scientific Workshop “Amphibians and Reptiles in Vietnam - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • PHUNG, TRUNG MY; MONA VAN SCHINGEN,, THOMAS ZIEGLER, & TRUONG QUANG NGUYEN 2014. A third new Cyrtodactylus (Squamata: Gekkonidae) from Ba Den Mountain, Tay Ninh Province, southern Vietnam. Zootaxa 3764 (3): 347–363 - get paper here
  • Rösler, Herbert; Thanh, Vu Ngoc; Truong, Nguyen Quang; Tri, Ngo Van & Ziegler, Thomas 2008. A new Cyrtodactylus (Squamata: Gekkonidae) from Central Vietnam. Hamadryad 33 (1): 48 – 63 - get paper here
  • SCHNEIDER, NICOLE; TRUNG MY PHUNG, MINH DUC LE,TRUONG QUANG NGUYEN, & THOMAS ZIEGLER 2014. A new Cyrtodactylus (Squamata: Gekkonidae) from Khanh Hoa Province, southern Vietnam. Zootaxa 3785 (4): 518–532 - get paper here
  • Stuart, B.L., Rowley, J.J.L., Neang, T., Emmett, D.A. & Sitha, Som 2010. Significant new records of amphibians and reptiles from Virachey National Park, northeastern Cambodia. Cambodian Journal of Natural History 2010(1): 38–47 - get paper here
  • Tri, Ngo Van & Onn, Chan Kinn 2010. A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa province, Southern Vietnam. Zootaxa 2504: 47–60 - get paper here
  • Ziegler, T.; ROMAN NAZAROV, NIKOLAI ORLOV, TRUONG QUANG NGUYEN, THANH NGOC VU, KIEN NGOC DANG, TRI HUY DINH, ANDREAS SCHMITZ 2010. A third new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha-Ke Bang National Park, Truong Son Range, Vietnam. Zootaxa 2413: 20–36 - get paper here
  • Ziegler, T.; TRUNG MY PHUNG, MINH DUC LE, Truong QUang Nguyen 2013. A new Cyrtodactylus (Squamata: Gekkonidae) from Phu Yen Province, southern Vietnam. Zootaxa 3686 (4): 432–446 - get paper here
 
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