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Cyrtodactylus timur GRISMER, WOOD, ANUAR, QUAH, MUIN, MOHAMED, ONN, SUMARLI, LOREDO & HEINZ, 2014

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Banjaran Timur Bent-toed Gecko 
SynonymCyrtodactylus timur GRISMER, WOOD, ANUAR, QUAH, MUIN, MOHAMED, ONN, SUMARLI, LOREDO & HEINZ 2014 
DistributionNE Peninsular Malaysia

Type locality: Punca Air, Gunung Tebu, Terengganu, Peninsular Malaysia (5° 36.11 N 102° 36.19 E; at 650 m elevation).  
Reproductionoviparous (manual and phylogenetic imputation, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 11207, adult male, collected on 1 July 2013 by Shahrul Anuar at 2350 hrs. Paratypes. LSUHC 11183–85 have the same collection data as the holotype. LSUHC 11288 has the same collection data as the holotype except that it was collected on 2 July 2013. 
DiagnosisDiagnosis: Cyrtodactylus timur sp. nov. can be differentiated from all other species of Cyrtodactylus by having 10–12 supralabials; 8–10 infralabials; weak body tuberculation; no tubercles on ventral surface of forelimbs, gular region, or in ventrolateral body folds; 38–43 paravertebral tubercles; 21–24 longitudinal tubercle rows; 31–40 ventral scales; 21–25 subdigital lamellae on fourth toe; 21 or 22 femoroprecloacal pores; deep precloacal groove; four dark dorsal body bands; body band/interspace ratio 1.00–1.25; no scattered white tubercles on dorsum; 8–10 dark caudal bands on original tail; white caudal bands nearly immaculate; and a maximum SVL of 120.5 mm. These characters are scored across all species of the C. pulchellus complex in Table 5 (Grismer et al. 2014).
 
CommentHAbitat: dipterocarp forest in the vicinity of a rocky stream; most common on the granite boulders along the stream but others were seen on the bases of trees and one individual was observed on an earthen bank.

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet “timur” is an invariable noun in apposition in reference to this species being found in the Banjaran Timur of northeastern Peninsular Malaysia. “Timur” means east in the Malay language and is in reference to the mountain range being on the east coast of Peninsular Malaysia. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, SHAHRUL ANUAR, EVAN S. H. QUAH, MOHD ABDUL MUIN, MAKETAB MOHAMED, CHAN KIN ONN, ALEXANDRA X. SUMARLI, ARIEL I. LOREDO, HEATHER M. HEINZ 2014. The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia. Zootaxa 3786 (3): 359–381 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Termprayoon, K., Rujirawan, A., Grismer, L. L., Wood Jr, P. L., & Aowphol, A. 2021. Taxonomic reassessment and phylogenetic placement of Cyrtodactylus phuketensis (Reptilia, Gekkonidae) based on morphological and molecular evidence. ZooKeys, 1040, 91 - get paper here
  • WOOD JR, PERRY L.; L. LEE GRISMER, MOHD ABDUL MUIN, SHAHRUL ANUAR, JAMIE R. OAKS 2020. A new potentially endangered limestone-associated Bent-toed Gecko of the Cyrtodactylus pulchellus (Squamata: Gekkonidae) complex from northern Peninsular Malaysia. Zootaxa 4751 (3): 437–460 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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