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Cyrtodactylus atremus KRAUS & WEIJOLA, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus atremus KRAUS & WEIJOLA 2019
Cyrtodactylus sp. ‘Karkar’ — TALLOWIN et al. 2018 
DistributionPapua New Guinea (Madang)

Type locality: above Mom, 4.6248°S, 145.9378°E, 500–645 m a.s.l., Karkar Island, Madang Province, Papua New Guinea  
Reproduction 
TypesHolotype: UMMZ 247747 (field tag FK 17685), mature male collected by V. Weijola, 26 February 2018.
Paratypes (n = 8). Male: Papua New Guinea: Madang Province: Karkar Island: same data as holotype (UMMZ 247746); Females: Papua New Guinea: Madang Province: Karkar Island: same data as holotype (UMMZ 247748– 51), Mom, 4.61°S, 145.92°E, 290 m a.s.l. (AMS R24744, R28901), Kevasob, 4.7°S, 145.9°E, 130 m a.s.l. (AMS R124604). 
DiagnosisDiagnosis. A moderately large (adult SVL = 108–118 mm) species of Cyrtodactylus with a small patch of somewhat enlarged precloacal scales (ca. 2–3 times size of nearby femoral scales), no enlarged femoral scales, no precloacal/femoral pores in males, and having a highly tuberculate dorsum, 22–24 rows of dorsal tubercles between the lateral skin folds, dorsal tubercles 3–5 times size of adjacent granules, 24–31 scale rows below the fourth toe, 9–11 supralabials to the center of eye, 10–13 infralabials to rictus, 40–47 mid-belly scale rows, 5–6 dark dorsal crossbands to the base of the tail, uniformly brown top of head, and lips dirty white with some brown stippling.

Comparisons with other species. The absence of enlarged femoral scales and precloacal/femoral pores in males distinguishes the new species from all other Papuan Cyrtodactylus except C. sermowaiensis (Kraus 2008). From that species, C. atremus differs in its larger size (adult SVL = 108–118 mm versus 85–94 mm in C. sermowaiensis; adult mass = 24.3–25.8 g versus 10.3–15.8 g in C. sermowaiensis); more tuberculate dorsum, limbs, and head (cf. Fig. 1 versus Fig. 2), with tubercles 3–5 times size of adjacent granules (versus 2–3 times in C. sermowaiensis); greater average number of rows of dorsal tubercles between the lateral skin folds (mean = 22.6, range = 22–24, SD = 0.799 versus mean = 20.3, range = 18–25, SD = 0.843 in C. sermowaiensis); fewer dark dorsal crossbands from head to tail base (mean = 5.8, range = 5–6, SD = 0.167 versus mean = 8.0, range = 7–9, SD = 0.365 in C. sermowaiensis); top of head uniformly brown (cf. Fig. 1A, 3A versus heavily patterned with dark-brown blotches, cf. Fig. 2A, 3B, C, D in C. sermowaiensis); and lips dirty white with some brown stippling (cf. Fig. 1B, 3A versus dark brown with few white or yellow spots, cf. Fig. 2B, 3B, C, D in C. sermowaiensis). 
Comment 
EtymologyThe name is a Latinized masculine compound adjective formed from the Greek a, meaning without, and trema, meaning hole. It refers to the lack of precloacal / femoral pores in males of this species. 
References
  • KRAUS, FRED; VALTER WEIJOLA 2019. New species of Cyrtodactylus (Squamata: Gekkonidae) from Karkar Island, Papua New Guinea. Zootaxa 4695 (6): 529–540 - get paper here
  • Tallowin, O. J. S., Tamar, K., Meiri, S., Allison, A., Kraus, F., Richards, S. J., & Oliver, P. M. 2018. Early insularity and subsequent mountain uplift were complementary drivers of diversification in a Melanesian lizard radiation (Gekkonidae: Cyrtodactylus). Molecular Phylogenetics and Evolution 125: 29–39 - get paper here
 
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