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Cyrtodactylus crustulus OLIVER, HARTMAN, TURNER, WILDE, AUSTIN & RICHARDS, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Manus bent-toed gecko 
SynonymCyrtodactylus crustulus OLIVER, HARTMAN, TURNER, WILDE, AUSTIN & RICHARDS 2020
Cyrtodactylus sp. ‘Manus’ — TALLOWIN et al. 2018: 33 
DistributionPapua New Guinea (Manus Island)

Type locality: Papua New Guinea, Manus Province, Manus Island, “Yeri camp” (-2.0010, 146.8190),  
TypesHolotype: SAMA R71122 (Field number SJR15091), adult male with original tail, with alcohol-preserved tissue for genetic analysis stored in the Australian Biological Tissues Collection (ABTC 136633), collected by Stephen Richards on 12 October 2014.
Paratypes. (n=4). SAMA R71119 (SJR15075) subadult, SAMA R71120–1 (SJR15076–7), SJR 15092 (to be repatriated to Papua New Guinea) adult males, all with same locality and collector information as holotype, collected between 11–12 October 2014. 
DiagnosisDiagnosis: A species of Cyrtodactylus that can be distinguished from all other species of Papuan Cyrtodactylus by the following unique combination of character states: moderate adult size (SVL 93–102 mm); continuous medial row of transversely enlarged subcaudal scales absent; enlarged femoral scales absent; precloacal and femoral pores in males absent; irregularly spaced dark-brown dorsal bands or transverse series of blotches on torso in series of 5–6 (not including nuchal band) with alternating indistinct lighter-brown interstitial bands; dorsal and lateral tubercles present on proximal and distal segments of all limbs; dorsal tubercles at mid-point of body between lateral skin folds in 18–19 rows; tubercles across supra- and interorbital regions usually >10; majority (80–90%) of tubercles within dark-brown dorsal bands light brown and contrasting against surrounding pigmentation; lateral region of head and torso with extensive yellowish spots in life; pale regions on dorsal surfaces of neck and posterior head with indistinct aggregations of light-brown subcircular markings; and nuchal band with regularly scalloped anterior and posterior edges. 
EtymologyCrustulum “cake”. Latin (masculine) meaning cake or pastry; used in reference to the small cake-shaped markings on the posterior dorsal region of the head and similar-shaped indentations along the edges of the nuchal band. 
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • OLIVER, PAUL M.; RYAN HARTMAN, CAMERON D. TURNER, TAYLOR A. WILDE, CHRISTOPHER C. AUSTIN, STEPHEN J. RICHARDS 2020. A new species of Cyrtodactylus Gray (Gekkonidae: Squamata) from Manus Island, and extended description and range extension for Cyrtodactylus sermowaiensis (De Rooij). Zootaxa 4728 (3): 341–356 - get paper here
  • Tallowin, O. J. S., Tamar, K., Meiri, S., Allison, A., Kraus, F., Richards, S. J., & Oliver, P. M. 2018. Early insularity and subsequent mountain uplift were complementary drivers of diversification in a Melanesian lizard radiation (Gekkonidae: Cyrtodactylus). Molecular Phylogenetics and Evolution 125: 29–39 - get paper here
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