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Cyrtodactylus dianxiensis LIU & RAO, 2021

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Western Yunnan Bent-toed Gecko
Chinese: 滇西裸趾虎’ (Diān Xī Luǒ Zhǐ Hǔ) 
SynonymCyrtodactylus dianxiensis LIU & RAO 2021
Cyrtodactylus khasiensis — YANG & RAO 2008 
DistributionChina (Yingjiang County and Longchuan County in Tongbiguan Nature Reserve, Dehong Autonomous Prefecture, Western Yunnan)

Type locality: Tongbiguan Township, Yingjiang County, Dehong Autonomous Prefecture, Yunnan Province, China (24° 36′ 51″ N, 97° 35′ 2″ E, at an elevation of 1170 m asl).  
Reproduction 
TypesHolotype: KIZ L2019044, adult male, collected by Shuo Liu on 3 September 2018.
Paratypes: KIZ 2003170, adult male, collected on 14 May 2003. 
DiagnosisDiagnosis: Crytodactylus dianxiensis can be diagnosed from other members of the C. gansi group by the following combination of characters: body size moderate (SVL 73.8–79.9 mm in adults); 11–12 supralabials; 9–11 infralabials; 18–19 longitudinal rows of rounded, conical, feebly keeled dorsal tubercles; 31–32 paravertebral tubercles; 37–41 ventral scales between ventrolateral folds; no precloacal groove; 7–8 precloacals in males, no femoral pores; 16 total subdigital lamellae beneath finger IV of manus, 19–20 total subdigital lamellae beneath toe IV of pes; dorsal pattern of indistinct light and dark blotches forming longitudinal markings on neck and forebody; tail with alternating dark and light bands. (Liu et al. 2021).

Comparisons: Cyrtodactylus dianxiensis can be distinguished from all other species in the C. gansi group. It differs from C. guwahatiensis Agarwal, Mahony, Giri, Chaitanya, and Bauer, C. tamaiensis (Smith), and C. tripuraensis Agarwal, Mahony, Giri, Chaitanya, and Bauer in lacking, as opposed to having femoral pores. It differs from C. aunglini, C. ayeyarwadyensis Bauer, C. gansi, C. guwahatiensis, C. jaintianensis Agarwal, Mahony, Giri, Chaitanya, and Bauer, C. kazirangaensis Agarwal, Mahony, Giri, Chaitanya, and Bauer, C. khasiensis, C. septentrionalis Agarwal, Mahony, Giri, Chaitanya, and Bauer, C. tamaiensis, and C. tripuraensis in having fewer precloacal or precloacofemoral pores. It differs from C. brevidactylus Bauer and C. gansi in having a distinct ventrolateral fold. It differs from C. ayeyarwadyensis, C. guwahatiensis, C. septentrionalis, C. tamaiensis, and C. tripuraensis in lacking, as opposed to having enlarged femoral scales. It differs from C. gansi in lacking, as opposed to having, a precloacal groove in males. It differs from C. chrysopylos Bauer in lacking band on nape and bands on back, as opposed to having. It differs from C. montanus Agarwal, Mahony, Giri, Chaitanya, and Bauer in having fewer longitudinal rows of dorsal tubercles and fewer paravertebral tubercles. It differs from C. myaleiktaung Grismer, Wood, Thura, Win, Grismer, Trueblood, and Quah in having enlarged post-precloacal scales, as opposed to lacking; lacking band on nape, as opposed to having; top of head with numerous brown spots, as opposed, top of head generally unicolour. Cyrtodactylus dianxiensis most closely resembles C. cayuensis Li, C. mandalayensis Mahony, and C. nagalandensis but is distinguishable from them. It differs from C. cayuensis in having 37–41 ventral scales between ventrolateral folds, as opposed to 28–34; a W-shaped black stripe on the occiput, as opposed to no W-shaped black stripe on the occiput; black caudal bands that gradually widen, as opposed to the black caudal bands of almost the same width. It can be distinguished from C. mandalayensis by its relative length of digits of the manus: IV > III > V > II > I, as opposed to IV > III > II > V > I; the scales between the precloacal pores and vent are obviously enlarged, as opposed to slightly enlarged; precloacal pores in an almost continuous angular series with exception of a single non-pore-bearing scale in the middle, as opposed to precloacal pores being continuous; no additional enlarged pore-bearing scale situated immediately posterior to the angle of the precloacal pore series, as opposed to an additional enlarged porebearing scale situated immediately posterior to the angle of the continuous precloacal pore series; a W-shaped black stripe on the occiput, as opposed to no W-shaped black stripe on the occiput. It differs from C. nagalandensis by its relative length of digits of the pes: IV > V > III > II > I, as opposed to IV > III >V > II > I; total subdigital lamellae on the fourth finger 16, as opposed to 12–14; total subdigital lamellae on the fourth toe 19–20, as opposed to 16; paravertebral tubercles 31–32, as opposed to 34–35. (Liu et al. 2021).

Color in life: Dorsal colour of head, body, limbs, and tail yellowish white; top of head with numerous brown spots; upper eyelid light blue, the edge of the upper eyelid yellow; a W-shaped black stripe on the occiput; labials brownish grey with many small white spots; an indistinct brownish-black postorbital streak that extends above ear opening; neck and back with black spots that form longitudinal markings on neck and forebody; limbs with indistinct brown bars and spots; 10 black and nine white caudal bands, black bands wider than white bands; ventral surfaces of body and limbs white, ventral surface of head pinkish-white, ventral surface of tail checkered with yellow and white; iris blueish grey with dark reticulations. (Liu et al. 2021). 
Comment 
Etymology‘Dianxi’ in Chinese language is a noun and means Western Yunnan, the specific epithet refers to the distribution of the new species in the westernmost part of Yunnan Province. 
References
  • Liu, Shuo & Dingqi Rao 2021. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Western Yunnan, China. Journal of Natural History, 55: 713-731 - get paper here
  • Yang DT, Rao DQ. 2008. Amphibia and Reptilia of Yunnan. Kunming: Yunnan Publishing Group Corporation, 1-411. (in Chinese)
 
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