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Cyrtodactylus fumosus (MÜLLER, 1895)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common Names 
SynonymGymnodactylus fumosus MÜLLER 1894: 833
Gymnodactylus fumosus — DE ROOIJ 1915: 16 (partim)
Cyrtodactylus fumosus — UNDERWOOD 1954: 475
Gymnodactylus fumosus — WERMUTH 1965: 52
Cyrtodactylus fumosus — GRISMER & LEONG 2005
Cyrtodactylus fumosus — RÖSLER & GLAW 2008
Cyrtodactylus fumosus — MECKE et al. 2016 
DistributionIndonesia (restricted to Sulawesi [= Celebes] by Mecke et al. 2016)

Type locality: “Boelawa-Gebirge (1200 m hoch [= elevation]), Celebes” (= Huidu Matabulawa), corrected to “Bone Mountains” (= Pegunungan Bone, North Sulawesi Province, Indonesia) by Boulenger (1897).  
TypesHolotype: NMBA (= NHMB) 2662 
DiagnosisHead large, depressed, oviform; snout longer than the diameter of the orbit, which nearly equals its distance from the ear-opening; forehead concave; ear-opening horizontal ) or oval and oblique, one half to two-thirds the diameter of the eye. Head granular, the granules slightly enlarged on the snout, with very small tubercles on the temple. Rostral nearly twice as broad as high, subquadrangular, with median cleft above, in contact with two or three small shields at its upper border; nostril bordered by the rostral and three scales, sometimes also by the first upper labial. Nine or ten upper and eight or nine lower labials; mental triangular, more broad than long; a pair of large chin-shields forming a suture behind the mental, with one or two much smaller shields on each side. Body covered above with small granules and few, widely scattered, roundish, flat and smooth or slightly keeled, small tubercles. A distinct lateral fold. Ventral scales small, cycloid, imbricate, smooth, 35 40 across the middle of the belly. Male with a pubic groove and a continuous series of 42 52 pores, pierced in enlarged scales, which are visible in the female. Tail cylin- drical, tapering, covered with small flat scales, which are larger on the lower surface. Limbs long; digits strong, slightly de- pressed at the base, strongly compressed in the distal part; the basal phalanx with well developed transverse plates below (de Rooij 1915: 16).

Coloration: Grey, grey-brown, or pinkish-brown above, with blackish spots, which may form irregular transverse bands on the body; a dark streak from the eye to the shoulder. Tail with dark bands; reproduced tail longitudinally streaked with blackish. Lower parts whitish or brownish, uniform or with dark dots (de Rooij 1915: 16).

Size: Length of head and body 71 mm; tail 77 mm (de Rooij 1915: 16). 
CommentThis species has previously been considered as a synonym of C. marmoratus and is thus very similar to that species. Cyrtodactylus halmahericus has been removed from the synonymy of C. fumosus (or C. marmoratus).

Distribution: Reports from Java are members of the C. marmoratus complex, C. halmahericus (Halmahera, Soah, Konorah) 
  • Boulenger, G.A. 1897. A catalogue of the reptiles and batrachians of Celebes with special reference to the collections made by Drs P & F Sarasin in 1893-1896. Proc. Zool. Soc. London 1897: 193-237 [p228-237: batrachia] - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Grismer L. Lee & Leong, T.M. 2005. New Species of Cyrtodactylus (Squamata: Gekkonidae) from Southern Peninsular Malaysia. Journal of Herpetology 39 (4): 584-591 - get paper here
  • Grismer, L. Lee & Ahmad, N. 2008. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 1924: 53–68 - get paper here
  • Grismer, L. Lee; Chan K. Onn; Grismer, J.L.; Wood, P.L. & Belabut, D. 2008. Three new species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 1921: 1–23 - get paper here
  • Grismer, L. Lee; Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • HARTMANN, LUKAS; SVEN MECKE, MAX KIECKBUSCH, FELIX MADER, HINRICH KAISER 2016. A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895). Zootaxa 4067 (5): 552–568 - get paper here
  • Janiawati, Ida Ayu Ari; Mirza Dikari Kusrini, Ani Mardiastuti 2016. Structure and Composition of Reptile Communities in Human Modified Landscape in Gianyar Regency, Bali. HAYATI Journal of Biosciences, doi:10.1016/j.hjb.2016.06.006 - get paper here
  • Koch, A. 2012. Discovery, Diversity, and Distribution of the Amphibians and Reptiles of Sulawesi and its offshore islands. Edition Chimaira, 374 pp. [ISBN 978-3-89973-432-4] - get paper here
  • Kramer, Eugen 1979. Typenkatalog der Echsen im Naturhistorischen Museum Basel (BM), Stand 1978. [type catalogue] Revue Suisse de Zoologie 86 (1): 159-166 - get paper here
  • Mecke, Sven; Lukas Hartmann, Felix Mader, Max Kieckbusch, Hinrich Kaiser 2016. Redescription of Cyrtodactylus fumosus (Müller, 1895) (Reptilia: Squamata: Gekkonidae), with a revised identification key to the bent-toed geckos of Sulawesi. Acta Herpetologica 11 (2): 151-160 - get paper here
  • MECKE, SVEN; MAX KIECKBUSCH, LUKAS HARTMANN, HINRICH KAISER 2016. Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa 4175 (4): 353–365 - get paper here
  • Mertens, R. 1929. Zwei neue Haftzeher aus dem Indo—Australischen Archipel (Rept.). Senckenbergiana, 11:237-241.
  • Müller,F. 1894. Reptilien und Amphibien aus Celebes. Verh. Ges. Basel 10 (3): 825-843 [1895]
  • Müller,F. 1895. Reptilien und Amphibien aus Celebes. (II. Bericht). Verh. naturf. Ges. Basel 10: 862-869 - get paper here
  • OLIVER , PAUL; PAUL EDGAR, MUMPUNI, DJOKO T. ISKANDAR & RON LILLEY 2009. A new species of bent-toed gecko (Cyrtodactylus: Gekkonidae) from Seram Island, Indonesia. Zootaxa 2115: 47-55 - get paper here
  • Onn, Chan Kin & Norhayati Ahmad 2010. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389: 47–56 - get paper here
  • RIYANTO, AWAL; HELLEN KURNIATI, ANDREW ENGILIS, JR 2016. A new Bent-toed gecko (Squamata: Gekkonidae) from the Mekongga Mountains, South East Sulawesi, Indonesia. . Zootaxa 4109 (1): 059-072 - get paper here
  • RIYANTO, AWAL; L. LEE GRISMER, PERRY L. JR. WOOD 2015. The fourth Bent-toed Gecko of the genus Cyrtodactylus (Squamata: Gekkonidae) from Java, Indonesia. Zootaxa 4059 (2): 351–363 - get paper here
  • Rösler, H. & Glaw, F. 2008. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa 1729: 8–22 - get paper here
  • Underwood, Garth 1954. On the classifcation and evolution of geckos. Proc. Zool. Soc. London, 124 (3): 469-492
  • Welton, Luke J.; Cameron D. Siler, Arvin Diesmos, and Rafe M. Brown 2009. A NEW BENT-TOED GECKO (GENUS CYRTODACTYLUS) FROM SOUTHERN PALAWAN ISLAND, PHILIPPINES AND CLARIFICATION OF THE TAXONOMIC STATUS OF C. ANNULATUS. Herpetologica 65 (3): 328-343 - get paper here
  • Wermuth, H. 1965. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich (80): 1—246
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