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Cyrtodactylus guakanthanensis GRISMER, BELABUT, QUAH, ONN, WOOD & HASIM, 2014

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Gua Kanthan Bent-toed Gecko
Malay: Cicak Gua Kanthan 
SynonymCyrtodactylus guakanthanensis GRISMER, BELABUT, QUAH, ONN, WOOD & HASIM 2014 
DistributionPeninsular Malaysia (Perak)

Type locality: Gua Kanthan, Perak, Peninsular Malaysia (4°45.685’N, 101°07.322’E; 45 m elevation  
TypesHolotype: LSUHC 11322, adult male, collected by Daicus M. Belabut, Chan K. Onn, Evan S. H. Quah and L. Lee Grismer on 14 July 2013.
Paratypes. All paratypes (LSUHC 11321, 11323, 11325–36, 11339) bear the same data as the holotype. 
DiagnosisDiagnosis. Cyrtodactylus guakanthanensis sp. nov. can be distinguished from all other Sundaland species by having the following suite of character states: adult SVL 77.7–82.2 mm adult SVL; moderately sized, conical, weakly keeled, body tubercles; tubercles present on occiput, nape, and limbs, and extend posteriorly beyond base of tail; 37–44 ventral scales; no transversely enlarged, median, subcaudal scales; proximal subdigital lamellae transversely expanded; 19–21 subdigital lamellae on fourth toe; abrupt transition between posterior and ventral femoral scales; enlarged femoral scales; no femoral or precloacal pores; precloacal groove absent; wide, dark postorbital stripes from each eye extending posteriorly to the anterior margin of the shoulder region thence forming a transverse band across the anterior margin of the forelimbs; and body bearing five (rarely four) wide, bold, dark bands. The meristic characters are scored against C. sworderi, C. tebuensis and C. quadrivirgatus in Table 5 and against all other Sundaland species in Grismer et al. (2012b: Table 5).

Comparisons. Within the Cyrotdactylus sworderi complex, C. guakanthanensis is separated from C. quadrivirgatus by having an abrupt transition between small and large postfemoral scales and a banded dorsal pattern. From C. sworderi it differs in having fewer paravertebral tubercles (32–34 versus 36–38); lacking as opposed to having precloacal pores; having as opposed to lacking enlarged femoral scales; having wide as opposed to narrow whitish caudal bands (Fig. 1); and having a banded as opposed to spotted dorsal pattern (Fig. 1). It differs from C. tebuensis in potentially having few suprlabials (nine or 10 versus 11–13); having wide as opposed to narrow whitish caudal bands which are fewer in number (nine versus 10–14; Fig. 1); and having a banded versus a spotted or striped dorsal pattern (Fig. 1). These differences are compared between all members of the C. sworderi complex in Table 5 (GRISMER et al. 2014). 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet guakanthanensis is an adjective and refers to the type locality Gua Kanthan (=Kanthan Cave) within the Gunung Kanthan limestone forest. 
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; DAICUS M. BELABUT,, EVAN S. H. QUAH, CHAN KIN ONN, PERRY L. WOOD, JR. & ROSLI HASIM 2014. A new species of karst forest-adapted Bent-toed Gecko (genus Cyrtodactylus Gray, 1827) belonging to the C. sworderi complex from a threatened karst forest in Perak, Peninsular Malaysia. Zootaxa 3755 (5): 434–446 - get paper here
  • Grismer, L.L.; P. L. WOOD, JR., SHAHRUL ANUAR, H. R. DAVIS, A. J. COBOS & M. L. MURDOCH 2016. A new species of karst forest Bent-toed Gecko (genus Cyrtodactylus Gray) not yet threatened by foreign cement companies and a summary of Peninsular Malaysia’s endemic karst forest herpetofauna and the need for its conservation. Zootaxa 4061 (1): 001–017 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR, SHAHRUL ANUAR, MARTA S. GRISMER, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MOHD ABDUL MUIN, HAYDEN R. DAVIS, CÉSAR AGUILAR, RANDY KLABACKA, ANTHONY J. COBOS, ANCHALEE AOWPHOL, JACK W. SITES, JR 2016. Two new Bent-toed Geckos of the Cyrtodactylus pulchellus complex from Peninsular Malaysia and multiple instances of convergent adaptation to limestone forest ecosystems. Zootaxa 4105 (5): 401-429 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
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