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Cyrtodactylus lenya MULCAHY, THURA & ZUG, 2017

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Lenya Banded Bent-toed Gecko 
SynonymCyrtodactylus lenya MULCAHY, THURA & ZUG in CONNETTE et al. 2017 
DistributionMyanmar (Tanintharyi Region)

Type locality: Myanmar, Tanintharyi Region, in the proposed Lenya National Park Extension (11.60 ̊N, 99.33 ̊E)  
TypesHolotype: USNM 587788, adult female, collected by Daniel G. Mulcahy, 15 May 2016. Paratypes. USNM 587789, adult male collected 15 May 2016 by Daniel G. Mulcahy and Grant M. Connette; CAS 260233, adult female collected 18 May 2016 by Grant M. Connette; both specimens within the vicinity of the holotype. 
DiagnosisDefinition. Midsize Cyrtodactylus of the C. oldhami species group, adults 73–74 mm SVL, not sexually dimorphic; 27% HeadL/SVL, 61–67% HeadW/HeadL, 41–46% HeadH/HeadL, 47% TrunkL/SVL, 16% ForeaL/SVL, 16% CrusL/SVL. Limbs slender, medium length digits of foreand hindfeet moderate (7–9% 4FingL/SVL;10–11% 4ToeL/SVL).
Dorsally head with granular scales, small tubercles in supratemporal area; 9 supralabials; 7–10 infralabials, one pair of enlarged postmentals. Dorsally trunk with 15–19 longitudinal rows of tubercles at midbody, 39–41 tubercles in paravertebral row; ventrolateral fold present but indistinct with intermittent large scales and without tubercles; 29 ventral trunk scales at midbody smooth, overlapping and 3–4X larger than dorsal granular scales. Tail with large tubercles dorsally on base, subcaudal scales distinctly enlarged, plate-like, and medially forming longitudinal row of rectangular scales. No precloacal groove or depression; distinctly enlarged row of precloacal and femoral scales but no precloacal or femoral pores; 2 cloacal spurs on each side. 5 proximal and 10–11 distal (15–16 total) 4FingLm; 6 proximal and 10–11 distal (17–18 total) 4ToeLm, basal distal lamellae on finger and toe paired.
Distinctly banded dorsally and laterally, broad, dark, two toned bands alternating with lighter interspaces; interspaces medium to light brown and half to two-thirds width of dark bands. Dark bands with narrow chocolate brown borders fore and aft of brown band (roughly anteroposterior width of interspace; five distinct dark bands, nuchal-cervical, scapular, and three trunk; less distinct sacral band followed by unicolor and equal-width brown and mediumbrown bands on tail. Nuchal-cervical band part of postorbital stripes of light dorsal stripe above broader brown stripe; this continuous supraorbital striping and nuchal-cervical band forming U-shaped nuchal collar. Dorsally head indistinctly mottled, although overall appearance nearly medium brown; loreal area medium brown; supralabial and lower temporal areas medium to light brown of interspaces; limbs medium brown dorsally; venter white. Preceding color description based on preserved specimens.

Morphological comparisons to malayan and peninsular Thailand Cyrtodactylus. Cyrtodactylus lenya and C. payarhtanensis appear to be members of the C. oldhami group of species that also includes C. oldhami, C. phetchaburiensis, C. peguensis, and C. tigroides. This group of mid-sized geckos (adults 50–80 mm SVL) is characterized by absence of a precloacal groove, presence of pubic patch of enlarged scales, no or few (0–8) precloacal pores, longitudinal row of enlarged precloacal and femoral scales, moderate to distinct ventrolateral trunk fold, enlarged rectangular subcaudal scales. Dorsal pattern is variable but all share a broad nuchal collar with dark center narrowly edged by white; the collar is continuous (part of) with the postorbital striping. C. consobrinoides, C. lenya and C. tigroides are the only oldhami group members sharing a regular banded pattern (bands dark centers edged fore and aft in white); the dark bands are much narrower than lighter interspaces in C. consobrinoides, C. lenya has 15 or more rows of dorsal tubercles and C. tigroides 13 rows. C. payarhtaniensis and some C. phetchaburiensis share dorsal bands of irregular shape (bands often diagonally transverse and not white edged); former lacks precloacal pores, latter with 4–6 precloacal pores in males and sometimes showing longitudinal dorsal stripes [46]. C. payartanensis is most similar to C. variegatus in dorsal color pattern but lacks preanal and femoral pores. 
CommentHabitat: karst formation at elevations between 40 and 75 m. Surrounding areas were mature wet evergreen forest with the age of dominant dipterocarp trees estimated at 70–100+ years.

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific name refers to this species presence in the proposed Lenya National Park. The name is proposed as a noun in apposition 
  • Connette GM, Oswald P, Thura MK, LaJeunesse Connette KJ, Grindley ME, Songer M, et al. 2017. Rapid forest clearing in a Myanmar proposed national park threatens two newly discovered species of geckos (Gekkonidae: Cyrtodactylus). PLoS One 12 (4): e0174432 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Zug, G.R. & Mulcahy, D.G. 2019. Identification guide Amphibians and reptiles of South Tanintharyi. Fauna & Flora International, 101 pp.
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