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Cyrtodactylus macrotuberculatus GRISMER & AHMAD, 2008

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Tuberculate Bent-toed Gecko, Large Tubercled Bent-toed Gecko 
SynonymCyrtodactylus macrotuberculatus GRISMER & AHMAD 2008
Cyrtodactylus macrotuberculatus — GRISMER 2011
Cyrtodactylus macrotuberculatus — GRISMER & QUAH 2019 
DistributionW Malaysia (Pulau Langkawi, Kedah)

Type locality: beneath an overpass on the road to Gunung Raya (06°23.023 N, 99°49.126 E; 621 m elevation), Pulau Langkawi, Kedah, Peninsular Malaysia.  
TypesHolotype: ZRC 2.6754, adult male, collected by L. Lee Grismer, Perry L. Wood, Jr., Timothy A. Youmans, and Norsham Yaakob on 8 August 2005. Paratypes: ZRC 
DiagnosisDiagnosis. Cyrtodactylus macrotuberculatus is distinguished from all other Sunda Shelf species by having a maximum SVL of 120 mm; very large, trihedral, keeled, tubercles on body, limbs, and tail; tubercles present on occiput and top of head; no granular scales (only large tubercles) on dorsal surfaces of limbs; 19–22 ventral scales; transversely enlarged, median, subcaudal scales; proximal subdigital lamellae square; 21–23 subdigital lamellae on fourth toe; abrupt transition between posterior and ventral femoral scales; enlarged femoral and precloacal scales with a continuous series of 35–37 pore-bearing scales (femoero-precloacal scales); precloacal groove present, precloacal depression absent; no white recticulum on head; body and tail banded, limbs immaculate. The majority of these characters are summarized across all Sunda Shelf species in Table 1 in GRISMER & AHMAD 2008.

Diagnosis.—Adult males reaching 118.6 mm SVL, adult females reaching 113.4 mm SVL; 9–12 supralabials, 7–11 infralabials; tubercles of dorsum very large with no intervening smaller tubercles; tubercles on ventral surfaces of forelimbs, in gular region, and in ventrolateral body fold; 31–44 paravertebral tubercles; 19–27 longitudinal rows of dorsal tubercles; 17–28 rows of ventral scales; 20–24 subdigital lamellae on fourth toe; 28–40 femoro-precloacal pores in males; dorsum not bearing a scattered pattern of white tubercles; four body bands in adults lacking lightened centers and light colored tubercles; band to interspace ratio 1.00–1.50; 7–10 dark, caudal bands on original tail; white caudal bands in adults infused with dark pigment; and posterior portion of tail in hatchlings and juveniles banded not white. These characters are scored across all species of the Cyrtodactylus pulchellus complex in Table 6 [from GRISMER et al. 2012]. 
  • Chan Kin Onn & Norhayati Ahmad 2010. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389: 47–56 - get paper here
  • Goldberg, Stephen R. and L. Lee Grismer. 2014. Cyrtodactylus macrotuberculatus (tuberculate bent-toed gecko) reproduction. Herpetological Review 45 (2): 327 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L. Lee & Ahmad, N. 2008. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 1924: 53–68 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L. Lee; Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR, SHAHRUL ANUAR, MARTA S. GRISMER, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MOHD ABDUL MUIN, HAYDEN R. DAVIS, CÉSAR AGUILAR, RANDY KLABACKA, ANTHONY J. COBOS, ANCHALEE AOWPHOL, JACK W. SITES, JR 2016. Two new Bent-toed Geckos of the Cyrtodactylus pulchellus complex from Peninsular Malaysia and multiple instances of convergent adaptation to limestone forest ecosystems. Zootaxa 4105 (5): 401-429 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, MOHD. ABDUL MUIN, MONTRI SUMONTHA, NORHAYATI AHMAD, AARON M. BAUER, SANSAREEYA WANGKULANGKUL, JESSE L. GRISMER & OLIVIER S. G. PAUWELS 2012. A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa 3520: 1–55 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, SHAHRUL ANUAR, EVAN S. H. QUAH, MOHD ABDUL MUIN, MAKETAB MOHAMED, CHAN KIN ONN, ALEXANDRA X. SUMARLI, ARIEL I. LOREDO, HEATHER M. HEINZ 2014. The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia. Zootaxa 3786 (3): 359–381 - get paper here
  • ONN, CHAN KIN; L. LEE. GRISMER, DIONYSIUS S. SHARMA, DAICUS BELABUT, and NORHAYATI AHMA 2009. New herpetofaunal records for Perlis State Park and adjacent areas. Malayan Nature Journal 61 (4): 255 - 262
  • QUAH, EVAN S. H.; L. LEE GRISMER, PERRY L. WOOD, JR. & SHAHRUL ANUAR MOHD SAH 2019. The discovery and description of a new species of Bent-toed Gecko of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 4668 (1): 051–075 - get paper here
  • SUMONTHA, MONTRI; OLIVIER S.G. PAUWELS, KIRATI KUNYA, AWAT NITIKUL, PHAMON SAMPHANTHAMIT & L. LEE GRISMER 2012. A new forest-dwelling gecko from Phuket Island, Southern Thailand, related to Cyrtodactylus macrotuberculatus (Squamata: Gekkonidae). Zootaxa 3522: 61–72 - get paper here
  • WOOD JR, PERRY L.; L. LEE GRISMER, MOHD ABDUL MUIN, SHAHRUL ANUAR, JAMIE R. OAKS 2020. A new potentially endangered limestone-associated Bent-toed Gecko of the Cyrtodactylus pulchellus (Squamata: Gekkonidae) complex from northern Peninsular Malaysia. Zootaxa 4751 (3): 437–460 - get paper here
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