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Cyrtodactylus majulah GRISMER, WOOD & LIM, 2012

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common Names 
SynonymCyrtodactylus majulah GRISMER, WOOD & LIM 2012
Gymnodactylus marmoratus – SWORDER 1925: 63 (not of KUHL)
Gymnodactylus marmoratus – SMITH 1930: 12
Cyrtodactylus quadrivirgatus – LIM & LIM 1992: 121 (not of TAYLOR)
Cyrtodactylus quadrivirgatus – BAKER & LIM 2008: 80
Cyrtodactylus quadrivirgatus – DRING 1979: 228 (in part)
Cyrtodactylus quadrivirgatus – DENZER & MANTHEY 1991: 314
Cyrtodactylus quadrivirgatus – CHAN-ARD et al., 1999: 24
Cyrtodactylus quadrivirgatus – MANTHEY & GROSSMANN 1997: 227 
DistributionSingapore (Central Catchment Nature Reserve); Indonesia (Pulau Bintan in the Riau Archipelago)

Type locality: Singapore: Central Catchment Nature Reserve, Nee Soon Swamp-forest (1°22'48.99"N, 103°49'05.39"E; 51 m elevation).  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZRC 2.6950, adult male, 67.9 mm SVL; coll: L. Grismer, E. Quah, K. O. Chan, A. Figueroa & A. Devan-Song, 22 Jun.2011. Paratypes. — Adult male: 68.0 mm SVL (ZRC 2.6951); adult male: 62.7 mm SVL (ZRC 2.6952); adult male: 67.2 mm SVL (LSUHC 10458); adult male: 67.2 mm SVL (ZRC 2.6953); collection data as per holotype. 
DiagnosisDiagnosis: — Cyrtodactylus majulah new species is differentiated from its Sundaland congeners by having the unique combination of a maximum SVL of 68.0 mm; eight or nine supralabials; seven or eight infralabials; strongly tuberculated body and limbs; 39–46 paravertebral tubercles; 43–53 ventral scales; no abrupt contact of large and small postfemoral scales; no enlarged femoral scales; 20–23 subdigital lamellae; no femoral pores; no deep precloacal groove; enlarged precloacal scales; 7–11 precloacal pores; no enlarged median subcaudals; tubercles on the anterior portion of the tail; no reticulated pattern on head; having a blotched dorsal pattern; and lacking the paired, dark, semi-lunar shaped blotches on the upper nape prominently outlined in white. These and other characters are scored across all Sundaland species (Table 4). 
CommentSynonymy: after GRISMER et al. 2012. 
EtymologyNamed after the Malay (Bahasa Melayu) word “majulah”, meaning to go forward, to progress. This is in allusion to the present rapid advancement of research in the taxonomy of Cyrtodactylus, in which many species have recently been formally named, and more are waiting to be described. 
  • FIGUEROA, ALEX; MARTYN E.Y. LOW, KELVIN K.P. LIM 2023. Singapore’s herpetofauna: updated and annotated checklist, history, conservation, and distribution. Zootaxa 5287 (1): 001–378 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L. Lee; Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Leo S, Suherman M, Permatasari A, Suganda D, Zulamri, Winarni NL. 2020. Herpetofauna diversity in Zamrud National Park, Indonesia: baseline checklist for a Sumatra peat swamp forest ecosystem. Amphibian & Reptile Conservation 14(2) [General Section]: 250–263 (e249) - get paper here
  • RIYANTO, AWAL; L. LEE GRISMER & PERRY L. WOOD, JR. 2015. Cyrtodactylus rosichonariefi sp. nov. (Squamata: Gekkonidae), a new swamp-dwelling bent-toed gecko from Bunguran Island (Great Natuna), Indonesia. Zootaxa 3964 (1): 114–124 [erratum in Zootaxa 3999 (4): 600] - get paper here
  • Teo, R.C.H. & N.J. Thomas 2019. Updated Inventory of Amphibians, Reptiles and Mammals of the Bukit Timah Nature Reserve, Singapore. Gardens’ Bulletin Singapore 71(Suppl. 1):145-183; doi: 10.26492/gbs71(suppl.1).2019-06 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
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