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Cyrtodactylus malayanus (DE ROOIJ, 1915)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Borneo Bow-fingered Gecko 
SynonymGymnodactylus malayanus DE ROOIJ 1915: 20
Cyrtodactylus baluensis — DE JONG 1928 (fide VAN ROOIJEN 2002)
Gymnodactylus malayanus — DAAN & HILLENIUS 1966: 130
Cyrtodactylus (Cyrtodactylus) malayanus — RÖSLER 2000: 66
Cyrtodactylus malayanus — GRISMER 2005
Cyrtodactylus malayanus — MECKE et al. 2016 
DistributionIndonesia (Borneo), Malaysia (Sarawak)

Type locality: Indonesia; exact locality unknown (fide DAAN & HILLENIUS 1966)  
TypesLectotype: ZMA.RENA 11309, adult male 
DiagnosisHead large, depressed, snout nearly one time and a half the diameter of the orbit, which equals its distance from the ear-opening; forehead and loreal region concave; ear-opening oval, oblique, one third the diameter of the eye. Head granular, with small tubercles on the occiput and temples, the granules largest on the snout. Rostral quadrangular, twice as broad as high, with median cleft above; nostril bordered by the rostral and three or four scales. Eight to ten upper and eight or nine lower labials; mental large, triangular; a large median pair of chin-shields (fig. 12 b], in contact behind the mental, followed by two smaller shields. Throat minutely granulate. Body long, covered above with very small granules and numerous small, round, keeled tubercles. Lateral fold distinct, with a few enlarged tubercles. Ventral scales small, cycloid, imbricate, 50-60 across the middle of the belly. Male with 9 (8) praeanal pores in an angular series (fig. 120) no pubic groove. Tail round, tapering, one and one third time the length of head and body, covered with small scales and rows of keeled, trihedral tubercles at the basal half; scales of the lower part enlarged. Limbs long; digits long, very little depressed at the base, strongly compressed in the distal part; basal part with small transverse plates below, narrower than the digit, rounded (fig. 12 c; de Rooij 1915: 20).

Coloration: Pale greyish-brown above, with dark brown transverse markings, separated on the vertebral line; flanks with a series of dark round spots, sometimes confluent with the transverse bands. Head with symmetrical dark spots, separated by greyish-white; a broad dark band from the eye to the nape, not meeting its fellow. Tail with ten dark brown transverse bands. Lower parts yellowish, each scale dotted with brown (de Rooij 1915: 20).

Size: Length of head and body 72 mm.; tail 95 mm (de Rooij 1915: 20). 
CommentDistribution: The occurrence of this species on Borneo is questionable fide MALKMUS et al. (2002).

Similar species: C. ingeri, C. consobrinus 
  • Ahmad, Norhayati; Eddie Ahmad, Mahathir Ratag, Edward Asrul Alimin Sinon, Brandon Don, Francer Francis, Muhammad Ridzuan Mahmod, Amshari Agimin, Daicus Belabut 2019. Amphibians and Reptiles of Imbak Canyon Study Centre and Batu Timbang Camp. Journal of Tropical Biology and Conservation 16: 25–33
  • Brongersma,L.D. 1934. Contributions to Indo-Australian herpetology. Zoologische Mededelingen 17: 161-251 - get paper here
  • Daan, S. & Hillenius,D. 1966. Catalogue of the type specimens of amphibians and reptiles in the Zoological Museum, Amsterdam. Beaufortia 13: 117-144
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. & Yaakob, N. 2007. Status of knowledge of the Malaysian herpetofauna. In Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. in: L. S. L. Chua, L. G. Kirton & L. G. Saw (eds.), Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31-81
  • Das, Indraneil D 2005. Bornean Geckos of the Genus Cyrtodactylus. Gekko 4 (2): 11-19
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Grismer L. Lee 2005. New species of Bent-toed Gecko (Cyrtodactylus Gray 1827) from Pulau Aur, Johor, West Malaysia. Journal of Herpetology 39 (3):424-432. - get paper here
  • Grismer L. Lee & Leong, T.M. 2005. New Species of Cyrtodactylus (Squamata: Gekkonidae) from Southern Peninsular Malaysia. Journal of Herpetology 39 (4): 584-591 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L. Lee & Ahmad, N. 2008. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 1924: 53–68 - get paper here
  • Grismer, L. Lee; Chan K. Onn; Grismer, J.L.; Wood, P.L. & Belabut, D. 2008. Three new species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 1921: 1–23 - get paper here
  • Grismer, L. Lee; Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Grismer, L.L.; Wood, P.L. & Youmans, T.M. 2007. Redescription of the gekkonid lizard Cyrtodactylus sworderi (SMITH 1925) from Southern Peninsular Malaysia. Hamadryad 31 (2): 250-257
  • HARTMANN, LUKAS; SVEN MECKE, MAX KIECKBUSCH, FELIX MADER, HINRICH KAISER 2016. A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895). Zootaxa 4067 (5): 552–568 - get paper here
  • Hikida T 1990. Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species. Japanese Journal of Herpetology 13 (3): 91-107 - get paper here
  • Jong, J.K. de 1928. Beiträge zur Kenntnis der Reptilienfauna von Niederländisch-Ost-Indien. Treubia 10:145-151 - get paper here
  • King, W. 1962. A new gekkonid lizard of the genus Cyrtodactylus from the Philippine Islands. Fieldiana, Zoology, 44:117—120. - get paper here
  • Linkem, Charles W.; Jimmy A. McGuire, Christopher J. Hayden, Mohammed Iqbal Setiadi, David P. Bickford, and Rafe M. Brown 2008. A New Species of Bent-Toe Gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica 64 (2): 224-234 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • MECKE, SVEN; MAX KIECKBUSCH, LUKAS HARTMANN, HINRICH KAISER 2016. Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa 4175 (4): 353–365 - get paper here
  • Onn, Chan Kin & Norhayati Ahmad 2010. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389: 47–56 - get paper here
  • Rooijen, J. van & van Rooijen, M. 2002. Eine erste Beobachtung eines mutmasslichen Cyrtodactylus malayanus (DE ROOIJ 1915) in Sarawak, Ost-Malaysia. Sauria 24 (2): 23-25 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, H. & Glaw, F. 2008. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa 1729: 8–22 - get paper here
  • Welton, L.J, Siler, C.D., Diesmos, A.C. & Brown, R.M. 2010. Phylogeny-based species delimitation of southern Philippines bent-toed geckos and a new species of Cyrtodactylus (Squamata: Gekkonidae) from western Mindanao and the Sulu Archipelago. Zootaxa 2390: 49–68 - get paper here
  • Welton, Luke J.; Cameron D. Siler, Arvin Diesmos, and Rafe M. Brown 2009. A NEW BENT-TOED GECKO (GENUS CYRTODACTYLUS) FROM SOUTHERN PALAWAN ISLAND, PHILIPPINES AND CLARIFICATION OF THE TAXONOMIC STATUS OF C. ANNULATUS. Herpetologica 65 (3): 328-343 - get paper here
  • Youmans, Timothy M. & L. Lee Grismer 2006. A new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History 10 (1): 61-70
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