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Cyrtodactylus ngoiensis SCHNEIDER, LUU, SITTHIVONG, TEYNIÉ, LE, NGUYEN & ZIEGLER, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Ngoi Bent-toed Gecko 
SynonymCyrtodactylus ngoiensis SCHNEIDER, LUU, SITTHIVONG, TEYNIÉ, LE, NGUYEN & ZIEGLER 2020 
DistributionLaos (Luang Prabang)

Type locality: Luang Prabang Province, Ngoi District (elevation of 420 m a.s.l.).  
Reproduction 
TypesHolotype. One adult male, IEBR 4548 (Fig 4a), collected by Alexandre Teynié in 2012.
Paratypes. Three adult females: ZFMK 102768, IEBR A.2013.110 (Fig. 4b), IEBR A.2013.111, collected by Truong Quang Nguyen and Alexandre Teynié in August 2013 from Luang Prabang Province, Ngoi District (at elevations of 402–427 m a.s.l.). 
DiagnosisCyrtodactylus ngoiensis sp. nov. differs from other closely related congeners by at least 11.6 % genetic divergence in the COI gene and can be diagnosed in morphology as follows: maximum SVL 95.3 mm; supralabials 6–9; infralabials 8–11; ventral scales 38–43; dorsal tubercles in 15–21 rows at midbody; enlarged femoral scales present; precloacal pores 7 in the male, 7 pitted scales in females; femoral pores 14 in the male, absent in females; five bright yellowish transverse dorsal bands; subcaudals enlarged. The new species is shown to be a member of the C. wayakonei species group, but morphologically resembles C. dumnuii from Thailand (Schneider et al. 2020).

Diagnosis. The new species can be distinguished from all its congeners on the basis of the following combination of characters: maximum SVL 95.3 mm; supralabials 6–9; infralabials 8–11; ventral scales 38–43; dorsal tubercles in 15–21 rows at midbody; enlarged femoral scales present; precloacal pores 7 in the male, 7 pitted scales in females; femoral pores 14 in the male, absent in females; five bright yellow transverse dorsal bands; subcaudals enlarged.
Comparisons. We compared the new species with its congeners from Laos (Table 2) and with other Cyrtodactylus from the Indochina region including Vietnam, Thailand and Cambodia (Table 3). Morphologically, Cyrtodactylus ngoiensis sp. nov. resembles the members of the C. wayakonei group due to the low amount of precloacal pores (Cyrtodactylus ngoiensis sp. nov.: PP in the male and (or pitted scales) in females: 7). Therefore, below we compared the new species with members of the C. wayakonei species group.
The new species has enlarged subcaudal scales and thus differs from the following species which are lacking enlarged subcaudals: C. bobrovi; C. otai; C. vilaphongi; C. wayakonei. The new species has femoral pores in males and thus differs from the following species which are lacking femoral pores in males: C. chauquangensis; C. cucphuongensis; Cyrtodactylus houaphanensis sp. nov.; C. martini; C. puhuensis; C. spelaeus; C. taybacensis. The new species has a contiguous series of precloacal-femoral pores in males and thus differs from the following species, which are separated by poreless scales: C. huongsonensis; C. sonlaensis; C. muangfuangensis. The new species differs from C. soni by having more femoral pores in males (14 vs. 5–8), and the presence of 7 precloacal pitted scales in females (vs. 7–8 pores).
Morphologically, Cyrtodactylus ngoiensis sp. nov. is most similar to C. dumnuii from Thailand but the new species can be distinguished from the latter by its larger size (maximum SVL 95.3 vs. 84.2), having more precloacal pores in males (7 vs. 5–6) and pitted precloacal scales in females (7 pitted scales vs. 0–7 pores) (Bauer et al. 2010). In addition, Cyrtodactylus ngoiensis sp. nov. has less pronounced dorsal tubercles than C. dumnuii and a different colour pattern (uniform brown bands in C. dumnuii versus thinner brown transverse bands edged in whitish yellow in Cyrtodactylus ngoiensis sp. nov.) (Schneider et al. 2020). 
Comment 
References
  • SCHNEIDER, NICOLE; VINH QUANG LUU, SALY SITTHIVONG, ALEXANDRE TEYNIÉ, MINH DUC LE, TRUONG QUANG NGUYEN, THOMAS ZIEGLER 2020. Two new species of Cyrtodactylus (Squamata: Gekkonidae) from northern Laos, including new finding and expanded diagnosis of C. bansocensis. Zootaxa 4822 (4): 503–530 - get paper here
 
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