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Cyrtodactylus petani RIYANTO, GRISMER & WOOD, 2015

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesIndonesian: Cicak Jari Lengkung Petani
E: Farmer’s Bent-toed Gecko
E: Klakah bent-toed gecko [klakahensis]
G: Klakah Bogenfinger-Gecko [klakahensis] 
SynonymCyrtodactylus petani RIYANTO, GRISMER & WOOD 2015
Cyrtodactylus klakahensis HARTMANN, MECKE, KIECKBUSCH, MADER & KAISER 2016
Cyrtodactylus fumosus — MERTENS 1934: 689
Cyrtodactylus klakahensis — MECKE et al. 2016 
DistributionIndonesia (East Java)

Type locality: Purwodadi Botanical Garden, Purwodadi Village, Purwodadi Subdistrict, Pasuruan District, East Java Province, Indonesia (07°47’58.73” S; 112°44’13.73” E; 325 m elevation

klakahensis: Indonesia (Java); Type locality: Klakah, Lumajang Regency, Jawa Timur Province, Java, Indonesia  
TypesHolotype: MZB Lace.12899 (Field number AR5507), adult male, collected 18 October 2006 by Awal Riyanto and Mulyadi. Paratype. MZB.Lace.11706, 11707, 11708, 11709, 11710, 11711, 11712, 11713, 11714, 11715, adult males, Jeladri Village, Winangon Subdistrict, Pasuruan District, East Java Province, Indonesia (07o46’15.8” S; 112o58’00.5” E; 129 m asl), collected 4 March 2014 by Awal Riyanto; MZB.Lace.12143, adult male, Mliwang Village, Kerek Subdistrict, Tuban District, East Java Province, Indonesia (06°49’59.4”S, 111°51’59.8”E; 86 m asl) collected by Awal Riyanto and Wahyu Trilaksono; MZB.Lace.12898, an adult male, Porong River, Sidoarjo City, East Java Province, Indonesia (7°26'48.37"S; 112°28'08.70"E), collected 17 October 2006 by Awal Riyanto and Mulyadi; MZB.Lace.12900, adult male, Purwodadi Botanical Garden, Purwodadi Village, Pasuruan District, East Java Province, Indonesia (7°47'58"S; 112°44'13"E; 325 m asl), collected 18 October 2006 by Awal Riyanto and Mulyadi.
Holotype: SMF 22476 (Figures 1 & 3A–C; Table 2 in Hartmann et al. 2016), an adult male, collected in 1928 or 1929 by members of the Deutsche Limnologische Sunda-Expedition. Paratypes. SMF 22477 (Figure 2A; Table 2), an adult male; SMF 22478 (Figure 2B; Table 2), an adult female; and SMF 22479 (Figure 2C; Table 2), an unsexed juvenile. All specimens have the same collection information as the holotype [klakahensis] 
DiagnosisDiagnosis. Cyrtodactylus petani sp. nov. is distinguished from all Javan and Sundaic species by having a maximum SVL up to 57.2 mm; nine or 10 supralabials; seven or eight infralabials; strongly tuberculated body and limbs; 20–25 paravertebral tubercles; 30–35 ventral scales; enlarged femoral scales; enlarged precloacal scales; 17–18 subdigital lamellae on the fourth toe; 31–35 continuous precloacal and femoral pores in males, pores absent in females; precloacal groove absent, no enlarged median subcaudals; tubercles on anterior portion of tail; no reticulated pattern on head; paired dark blotches forming a V-shaped on occiput; blotched dorsal pattern; and no paired, dark, semi-lunar-shaped blotches on upper nape (Riyanto et al. 2015).

Diagnosis (klakahensis): Cyrtodactylus klakahensis can be distinguished from all other congeners occuring in the Greater Sunda Islands (including Sulawesi) and Lesser Sunda Islands by the following combination of characters: (1) a deep precloacal groove in males, (2) three rows of enlarged precloacofemoral scales, of which the third row bears 37–38 pores in males, (3) three or four rows of posterior precloacal scales, (4) raised and strongly keeled dorsal tubercles in 15–19 rows at midbody, (5) an indistinct lateral fold, (6) 17–20 subdigital lamellae under the 4th toe, and (7) subcaudal scales which are not transversely enlarged (Hartmann et al. 2016).

Comparisons. Cyrtodactylus petani sp.nov. differs from its sister species C. batucolus Grismer, Chan, Grismer, Wood & Belabut, 2008 by having fewer precloaco-femoral pores (31–35 versus 43–46) and a smaller maximum SVL (57.2 mm versus 75.2 mm). Cyrtodactylus petani sp. nov. differs from C. seribuatensis Grismer & Youmans, 2006 in having fewer precloaco-femoral pores (31–35 versus 40–43) and a smaller maximum SVL (57.2 mm versus 75 mm). The new species is distinguished from C. fumosus by lacking a precloacal groove, having fewer paravertebral tubercles (20–25 versus 30–33), fewer subdigital lamellae under the fourth toe (17–18 versus 22), fewer precloaco-femoral pores (31–35 versus 46–50) and a smaller maximum SVL (57.2 mm versus 75.2 mm). From C. marmoratus sensu stricto (lectotypes; see Rӧsler et al. 2007) it can differentiated by having tubercles in the ventrolateral body fold, fewer ventral scales (30–35 versus 38–47), fewer precloaco-femoral pores (31–35 versus 45–50) and fewer subdigital lamellae under the fourth toe (17–18 versus 20–24 lamellae). It can be differentiated from C. semiadii Riyanto, Bauer & Yudha, 2010 by having tuberculation on both forelimbs and hind limbs, presence of precloaco-femoral scales and fewer paravertebral tubercles (20–25 versus 37–40) (Riyanto et al. 2015) 
CommentSynonymy: riyanto et al. 2020 synonymized Cyrtodactylus klakahensis with C. petani. 
EtymologyThe specific epithet petani refers to the fact that the type series was collected on a farm. Petani means a farmer in the Indonesian Language and is here treated as a noun in apposition.

The specific epithet klakahensis is a Latinized, toponymic adjective referring to the type locality Klakah (Lumajang Regency, Jawa Timur Province, Java, Indonesia) of the new species. 
  • Amarasinghe, A. A.; Awal Riyanto, Mumpuni & Lee L. Grismer 2020. A NEW BENT-TOED GECKO SPECIES OF THE GENUS Cyrtodactylus GRAY, 1827 (SQUAMATA: GEKKONIDAE) FROM THE WEST BALI NATIONAL PARK, BALI, INDONESIA. TAPROBANICA 9 (1): 59–70, pls. 22–24. - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • HARTMANN, LUKAS; SVEN MECKE, MAX KIECKBUSCH, FELIX MADER, HINRICH KAISER 2016. A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895). Zootaxa 4067 (5): 552–568 - get paper here
  • MECKE, SVEN; MAX KIECKBUSCH, LUKAS HARTMANN, HINRICH KAISER 2016. Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa 4175 (4): 353–365 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Mertens, R. 1934. Die Amphibien und Reptilien der Deutschen Limnologischen Sunda-Expedition. In: Thienemann- Festband, A. (Ed.), Tropische Binnengewässer. Archiv für Hydrobiologie 40: 677–701
  • RIYANTO, AWAL; ACHMAD FARAJALLAH, AMIR HAMIDY, YULI SULISTYA FITRIANA, MISBAHUL MUNIR, NIA KURNIAWAN, ERIC N. SMITH 2020. Taxonomic evaluation of two similar bent-toed geckos Squamata: Gekkonidae: Cyrtodactylus Gray, 1827) from East Java, Indonesia. Zootaxa 4830 (1): 186–196 - get paper here
  • RIYANTO, AWAL; L. LEE GRISMER, PERRY L. JR. WOOD 2015. The fourth Bent-toed Gecko of the genus Cyrtodactylus (Squamata: Gekkonidae) from Java, Indonesia. Zootaxa 4059 (2): 351–363 - get paper here
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