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Cyrtodactylus thirakhupti PAUWELS, BAUER, SUMONTHA & CHANHOME, 2004

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Thirakhupt’s bent toed gecko
F: Cyrtodactyle de Thirakhupt
G: Thirakhupts Bogenfingergecko
Dutch: Kromvingergekko van Thirakhupt
Thai: Tuk kai Thirakhupt 
SynonymCyrtodactylus thirakhupti PAUWELS, BAUER, SUMONTHA & CHANHOME 2004
Cyrtodactylus thirakhupti — GRISMER 2005 
DistributionS Thailand (Surat Thani)

Type locality: Southern Thailand, Surat Thani Province, Thachana District, Tham Khao Sonk (= Cave of Sonk Mountain), 9°34’N 99°10’E, collected by Montri Sumontha, 27 June 2003.  
TypesHolotype: CUMZ (Chulalongkorn University Museum of Zoology) R 2003.120 
DiagnosisDiagnosis. — Cyrtodactylus thirakhupti can be distinguished from all congeners on the basis of the following combination of characters: body slender, limbs, digits and tail long, dorsal scalation with 14 rows of small keeled tubercles, 37-40 ventral scales across belly between weakly developed ventrolateral folds, no precloacal groove, precloacal pores and femoral pores absent in both sexes, femoral scales enlarged, median subcaudal scales enlarged to form broad transverse plates, and dorsal color pattern consisting of a series of dark-edged light bands. The absence of both precloacal and femoral pores distinguishes C. thirakhupti from all species except most members of the subgenus Geckoella: C. albofasciatus (Boulenger), C. collegalensis (Beddome), C. deccanensis (Günther), C. jeyporensis (Beddome), C. nebulosus (Beddome), and C. yakhuna (Deraniyagala), the Indonesian species C. jellesmae (Boulenger) and C. laevigatus Darevsky, C. paradoxus (Darevsky & Szczerbak) of Vietnam, an undescribed species from Laos (David et al., submitted), and C. sermowaiensis (de Rooij) of New Guinea. It may be distinguished from all Geckoella by its gracile body, long slender tail and long digits, from C. jellesmae, C. sermowaiensis, and the Laotian species by its enlarged series of femoral scales, and from the remaining species by its color pattern (dark-edged, light transverse bands versus dark spots in C. laevigatus and transverse bands interrupted by a narrow vertebral stripe in C. paradoxus). Some specimens of the Thai/ Myanmar species C. oldhami (Theobald) also lack pores (Smith 1935; Taylor 1963, Ulber 1993), but this species may be easily distinguished by its pattern of white spots and stripes. Although C. feae (Boulenger) of Myanmar was originally described as possessing pores, this has subsequently been called into question (see Bauer 2003). Regardless, it also differs markedly in color pattern from C. thirakhupti. Cyrtodactylus derongo Brown & Parker of New Guinea was described on the basis of females only and the condition of precloacal and femoral pores in males is unknown, but this species is much larger (to 112 mm SVL) than C. thirakhupti and differs markedly in color pattern and details of scalation. In general appearance the new species is most similar to the recently described C. sumonthai from Rayong Province, Thailand, which has only two tiny precloacal pores, which may be easily overlooked. However, this species lacks enlarged femoral scales and differs in details of color pattern. 
CommentDefinition. — A moderately sized Cyrtodactylus, snout-vent length to at least 80 mm; body slender, limbs and digits long, slender, original tail very long; one pair of enlarged postmental scales in broad contact with one another; dorsal scalation with 14 rows of keeled tubercles; approximately 37 ventral scales across belly between weakly-developed ventrolateral folds; no precloacal groove, no precloacal or femoral pores in either sex; eight broad basal lamellae and 12 narrow distal lamellae beneath 4th toe of pes; median subcaudal scales enlarged to form broad transverse plates; dorsal pattern of yellowish bands with very dark brown borders on a lighter brown background.

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyNamed after Prof. Dr Kumthorn Thirakhupt (Faculty of Science, Chulalongkorn University, Bangkok) in recognition of his contribution to Thai zoology. 
  • Areesirisuk P, Muangmai N, Kunya K, Singchat W, Sillapaprayoon S, Lapbenjakul S, Thapana W, Kantachumpoo A, Baicharoen S, Rerkamnuaychoke B, Peyachoknagul S, Han K, Srikulnath K. 2018. Characterization of five complete Cyrtodactylus mitogenome structures reveals low structural diversity and conservation of repeated sequences in the lineage. PeerJ 6: e6121 - get paper here
  • Bauer, A.M. 2003. Descriptions of seven new Cyrtodactylus (Squamata: Gekkonidae) with a key to the species of Myanmar (Burma). Proc. Cal. Acad. Sci. 54: 463-498 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Grismer L. Lee 2005. New species of Bent-toed Gecko (Cyrtodactylus Gray 1827) from Pulau Aur, Johor, West Malaysia. Journal of Herpetology 39 (3):424-432. - get paper here
  • Grismer L. Lee & Leong, T.M. 2005. New Species of Cyrtodactylus (Squamata: Gekkonidae) from Southern Peninsular Malaysia. Journal of Herpetology 39 (4): 584-591 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L.L.; Wood, P.L. & Youmans, T.M. 2007. Redescription of the gekkonid lizard Cyrtodactylus sworderi (SMITH 1925) from Southern Peninsular Malaysia. Hamadryad 31 (2): 250-257
  • Linkem, Charles W.; Jimmy A. McGuire, Christopher J. Hayden, Mohammed Iqbal Setiadi, David P. Bickford, and Rafe M. Brown 2008. A New Species of Bent-Toe Gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica 64 (2): 224-234 - get paper here
  • LUU, VINH QUANG; THOMAS CALAME, Truong QUang Nguyen, MICHAEL BONKOWSKI & THOMAS ZIEGLER 2015. A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa 4058 (3): 388–402 - get paper here
  • LUU, VINH QUANG; Truong QUang Nguyen, MINH DUC LE, MICHAEL BONKOWSKI, THOMAS ZIEGLER 2016. A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa 4079 (1): 087–102 - get paper here
  • LUU, VINH QUANG;THOMAS CALAME, MICHAEL BONKOWSKI, TRUONG QUANG NGUYEN & THOMAS ZIEGLER 2014. A new species of Cyrtodactylus (Squamata: Gekkonidae) from Khammouane Province, Laos. Zootaxa 3760 (1): 054–066 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • PAUWELS, O.S.G.; AARON M. BAUER; MONTRI SUMONTHA & LAWAN CHANHOME 2004. Cyrtodactylus thirakhupti (Squamata: Gekkonidae), a new cave-dwelling gecko from southern Thailand. Zootaxa 772: 1–11 - get paper here
  • Rösler, H. & Glaw, F. 2008. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa 1729: 8–22 - get paper here
  • Sumontha, M., Panitvong, N. & Deein, G. 2010. Cyrtodactylus auribalteatus (Squamata: Gekkonidae), a new cave- dwelling gecko from Phitsanulok Province, Thailand. Zootaxa 2370: 53–64 - get paper here
  • Tri, Ngo Van 2008. Two new cave-dwelling species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Southwestern Vietnam. Zootaxa 1909: 37–51 - get paper here
  • Ulber T 1993. Bemerkungen über cyrtodactyline Geckos aus Thailand nebst Beschreibungen von zwei neuen Arten (Reptilia: Gekkonidae). Mitt. Zool. Mus. Berlin 69(2) 1993: 187-200 - get paper here
  • Welton, Luke J.; Cameron D. Siler, Arvin Diesmos, and Rafe M. Brown 2009. A NEW BENT-TOED GECKO (GENUS CYRTODACTYLUS) FROM SOUTHERN PALAWAN ISLAND, PHILIPPINES AND CLARIFICATION OF THE TAXONOMIC STATUS OF C. ANNULATUS. Herpetologica 65 (3): 328-343 - get paper here
  • Youmans, Timothy M. & L. Lee Grismer 2006. A new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History 10 (1): 61-70
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