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Darevskia mixta (MÉHELY, 1909)

IUCN Red List - Darevskia mixta - Near Threatened, NT
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Higher TaxaLacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesG: Bastardeidechse 
SynonymLacerta mixta MÉHELY 1909: 581
Lacerta composita MÉHELY 1909 (nomen oblitum)
Lacerta saxicola mixta LANTZ & CYREN 1936
Lacerta (Zootoca) “mixta” — BISCHOFF 1978
Lacerta mixta — ENGELMANN et al 1993
Lacerta mixta — FU & MURPHY 1997
Darevskia mixta — ARRIBAS 1997
Lacerta mixta — FU, MURPHY & DAREVSKY 2000
Darevskia mixta — MURPHY et al. 2000
Archaeolacerta (Caucasilacerta) mixta — SINDACO et al. 2000
Lacerta mixta — SZCZERBAK 2003
Darevskia mixta — BISCHOFF 2007
Lacerta (Darevskia) mixta — SINDACO & JEREMČENKO 2008 
DistributionSW Republic of Georgia (Small Caucasus)

Type locality: Abastumani, Republic of Georgia  
Reproductionoviparous 
TypesLectotype: SMF 12087, designated by R. Mertens 1967. 
DiagnosisUnfortunately we had to temporarily remove additional information as this was scraped by multiple AI companies who sell that data to their customers. These details, e.g. detailed descriptions or comparisons (about 1988 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy partly after KHALIKOV & ANANJEVA (pers. comm.). This species is not listed in the Zoological Record (1909) where other species described by MÉHELY are listed.

Distribution: also in Turkey (fide web site Türkiye'de Kaç Tür KertenkeleYas¸ıyor), but not in Turkey fide Gabelaia et al. 2015. 
References
  • Abramjan, A., Arakelyan, M., & Frynta, D. 2020. Does reproductive mode affect sexually selected colouration? Evaluating UV-blue spots in parthenogenetic and bisexual lizards of the genus Darevskia. Current Zoology - get paper here
  • Bischoff, W. 1978. Beiträge zur Kenntnis der Echsen des Kaukasus. Salamandra 14 (4): 178-202 - get paper here
  • Bischoff, W. 2007. Auf der Suche nach Gebirgseidechsen in drei Erdteilen. Draco 7 (27): 54-73 - get paper here
  • Darevsky,I.S. 1957. Systematics and ecology of rock lizards (Lacerta saxicola Eversmann) in Armenia [in Russian]. Zool. sb. AN Armenia SSR 10: 27-57 - get paper here
  • Doronin, Igor V. 2018. The Taxonomic Status of Lacerta mixta Méhely, 1909 (Squamata: Sauria: Lacertidae). Russ. J. Herpetol. 25 (2): 143-146 - get paper here
  • Ebel, R., Müller, J., Ramm, T. et al. 2020. First evidence of convergent lifestyle signal in reptile skull roof microanatomy. BMC Biol 18, 185 (2020) - get paper here
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Freitas, S., Westram, A.M., Schwander, T., Arakelyan, M., Ilgaz, Ç., Kumlutas, Y., Harris, D.J., Carretero, M.A. and Butlin, R.K. 2022. Parthenogenesis in Darevskia lizards - a rare outcome of common hybridisation, not a common outcome of rare hybridisation Parthenogenesis, gene flow and hybridisation. Evolution 76 (5): 899–914<br /> - get paper here
  • Fu J.; Murphy R.W.; Darevsky I.S. 1999. Limited genetic variation in Lacerta mixta and its parthenogenetic daughter species: evidence from cytochrome b and ATPase 6 gene DNA sequences. Genetica 105 (3): 227-231
  • Fu, JINZHONG AND ROBERT W. MURPHY 1997. Toward the phylogeny of caucasian rock lizards: implications from mitochondrial DNA gene sequences (Reptilia: Lacertidae). Zoological Journal of the Linnean Society (1997), 121: 463–477. - get paper here
  • Gabelaia, Mariam, David Tarkhnishvili and Marine Murtskhvaladze. 2015. Phylogeography and morphological variation in a narrowly distributed Caucasian rock lizard, Darevskia mixta. Amphibia-Reptilia 36 (1): 45-54 - get paper here
  • Gabelaia, Mariam; Dominique Adriaens, David Tarkhnishvili 2017. Phylogenetic signals in scale shape in Caucasian rock lizards (Darevskia species). Zoologischer Anzeiger 268: 32–40 - get paper here
  • Grechko VV, Ciobanu DG, Darevsky IS, Kosushkin SA, Kramerov DA. 2006. Molecular evolution of satellite DNA repeats and speciation of lizards of the genus Darevskia (Sauria: Lacertidae). Genome 49(10):1297-307. - get paper here
  • Lantz,L.A. & Cyrén,O. 1936. Contribution à la connaissance de Lacerta saxicola EVERSMANN. Bull. Soc. zool. France (Paris) 61: 159-181 - get paper here
  • Méhely, Lajos von 1909. Materialien zu einer Systematik und Phylogenie der muralis-ähnlichen Lacerten. Annales historico-naturales Musei nationalis Hungarici, Budapest, 7 (2): 409-621 - get paper here
  • Murphy, R.W.; Fu,J.; MacCulloch, R.D.; Darevsky,I.S. & Kupriyanova,L.A. 2000. A fine line between sex and unisexuality: the phylogenetic constraints on parthenogenesis in lacertid lizards. Zool. J. Linnean Soc. 130: 527-549 - get paper here
  • Petrosyan, V., Osipov, F., Bobrov, V., Dergunova, N., Omelchenko, A., Varshavskiy, A., ... & Arakelyan, M. 2020. Species Distribution Models and Niche Partitioning among Unisexual Darevskia dahli and Its Parental Bisexual (D. portschinskii, D. mixta) Rock Lizards in the Caucasus. Mathematics, 8(8): 1329 - get paper here
  • Petrosyan, Varos; Fedor Osipov, Vladimir Bobrov, Natalia Dergunova, Evgeny Nazarenko, Andrey Omelchenko, Felix Danielyan & Marine Arakelyan 2019. Analysis of geographical distribution of the parthenogenetic rock lizard Darevskia armeniaca and its parental species (D. mixta, D. valentini) based on ecological modelling. Salamandra 55 (3): 173-190 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Alberto Venchi, Giuseppe M. Carpaneto, Marco A. Bologna 2000. The Reptiles of Anatolia: a Checklist and Zoogeographical analysis. Biogeographia, 21-2000: 441-554. - get paper here
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
  • Tarkhnishvili, D., Gabelaia, M. and Adriaens, D. 2020. Phenotypic divergence, convergence and evolution of Caucasian rock lizards (Darevskia). Biological Journal of the Linnean Society - get paper here
  • Tuniyev B.S., Tuniyev S.B., Avcı A., Ilgaz Ç. 2014. Herpetological studies in Eastern and North-Eastern Turkey. Current Studies in Herpetology 14 (1/2): 44–53. Резюме - get paper here
  • Tuniyev, B.S. 1990. On the Independence of the Colchis Center of Amphibian and Reptile Speciation. Asiatic Herpetological Research 3: 67-84 - get paper here
  • Uzzell, T. and Darevsky, I. S. 1973. Electrophoretic examination of Lacerta mixta - a possible hybrid species (Sauria, Lacertidae). Journal of Herpetology 7 (1): 11-15. - get paper here
  • Uzzell, T. and Darevsky, I. S. 1975. Biochemicai evidence for the hybrid origin of the parthenogenetic species of the Lacerta saxicola complex (Sauria: Lacertidae), with a dicussion of some ecological and evolutionary implications. Copeia 1975 (2): 204-222. - get paper here
 
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