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Darevskia obscura (LANTZ & CYRÉN, 1936)

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Higher TaxaLacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards)
SubspeciesDarevskia obscura bischoffi (BÖHME & BUDAK 1977)
Darevskia obscura macromaculata (DAREVSKY 1967)
Darevskia obscura obscura (LANTZ & CYRÉN 1936) 
Common NamesG: Grusinische Eidechse, Kielschwanz-Felseidechse
Russian: Грузинская ящерица 
SynonymLacerta saxicola obscura LANTZ & CYRÉN 1936: 177
Lacerta rudis obscura — LANGERWERF 1978
Lacerta (Darevskia) rudis obscura — SINDACO & JEREMČENKO 2008
Darevskia rudis obscura — ARRIBAS et al. 2013
Darevskia rudis obscura — TUNIYEV et al. 2014
Darevskia obscura — GABELAIA et al. 2018
Darevskia obscura — TARKNISHVILI et al. 2020
Darevskia obscura — ARRIBAS et al. 2022

Darevskia obscura bischoffi (BÖHME & BUDAK 1977)
Lacerta (Darevskia) rudis bischoffi — SINDACO & JEREMČENKO 2008
Darevskia rudis bischoffi — ARRIBAS et al. 2013
Darevskia rudis bischoffi — TUNIYEV et al. 2014
Darevskia obscura bischoffi — ARRIBAS et al. 2022

Darevskia obscura macromaculata (DAREVSKY 1967)
Lacerta (Darevskia) rudis macromaculata — SINDACO & JEREMČENKO 2008
Darevskia rudis macromaculata — ARRIBAS et al. 2013
Darevskia rudis macromaculata — TUNIYEV et al. 2014
Darevskia obscura macromaculata — ARRIBAS et al. 2022 
DistributionRepublic of Georgia, NE Turkey

bischoffi: NE Turkey; Type locality: Turkey, Artvin Province, 6 km to the southwest Arhavi, rocky exits along the automobile route, 41°19 ́15 ́ ́N 41°14 ́41 ́ ́E.

macromaculata: Republic of Georgia, NE Turkey; Type locality: Georgia, Samtskhe-Javakheti Region, vicinity of Akhalkalaki, about 1800 m asl, 41°23 ́N 43°29 ́E.

obscura: Republic of Georgia, NE Turkey; lectotype locality: Georgia, Samtskhe-Javakheti, Borjomi, road to Akhaltsikhe.  
Reproductionoviparous 
TypesLectotype: ZISP 17171.1, designated by Doronin 2017 (Fig. 7 in Doronin 2017), 30.05.1914. Coll.: L.A. Lantz; paralectotypes: BMNH 1918.11.21.5–7
Holotype: ZDEU 183/977, 05.1977. Coll.: A. Budak [bischoffi]
Holotype: ZISP 17940.6 (Figs. 8-9 in Doronin 2017), 24.06.1961. Coll.: I.S. Darevsky. [macromaculata] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (3292 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy partly after KHALIKOV & ANANJEVA (pers. comm.). RYABININA et al. (2002) question the separation of bischoffi and obscura. ARRIBAS et a. 2013 suggest that macromaculata is a synonym of obscura. However, Arribas et al. elevated obscura to full species status with two subspecies (bischoffi and macromaculata). We consider this elevation a bit preliminary, given that they apparently only used a fragment of cyt-b for their phylogenetic analysis.

Subspecies: Darevskia rudis bithynica (MÉHELY, 1909) has been elevated to species status, with D. r. tristis as subspecies. Gabelaia et al. 2018 elevated D. r. obscura to full species status, based on scalation characters. Lantz & Cyren 1936 did provide a diagnosis (in french) but did not illustrate the diagnostic characters.

Relationships: in Turkey, only minor genetic differences separated D. rudis and D. bithynica populations (Koç et al. 2021). 
References
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  • ARRIBAS, O., CANDAN, K., KORNILIOS, P., AYAZ, D., KUMLUTAŞ, Y., GÜL, S., ... & ILGAZ, Ç. 2022. Revising the taxonomy of Darevskia valentini (Boettger, 1892) and Darevskia rudis (Bedriaga, 1886)(Squamata, Lacertidae): a Morpho-Phylogenetic integrated study in a complex Anatolian scenario. Zootaxa 5224 (1): 1-68 - get paper here
  • ARRIBAS, OSCAR; ÇETİN ILGAZ, YUSUF KUMLUTAŞ, SALİH HAKAN DURMUŞ, AZİZ AVCI & NAZAN ÜZÜM 2013. External morphology and osteology of Darevskia rudis (Bedriaga, 1886), with a taxonomic revision of the Pontic and Small-Caucasus populations (Squamata: Lacertidae). Zootaxa 3626 (4): 401–428 - get paper here
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