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Dendragama boulengeri DORIA, 1888

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Boulenger's Tree Agama 
SynonymDendragama boulengeri DORIA 1888
Dendragama boulengeri — WERNER 1900: 484
Dendragama boulengeri — DE ROOIJ 1915: 118
Calotes (Pseudocalotes) boulengeri — MERTENS 1954: 185
Calotes boulengeri — WERMUTH 1967: 34
Calotes boulengeri — BRYGOO 1988: 9
Dendragama boulengeri — MANTHEY & GROSSMANN 1997: 167
Dendragama boulengeri — MANTHEY & SCHUSTER 1999: 51
Dendragama boulengeri — TEYNIÉ et al. 2010
Dendragama schneideri — HARVEY Et al. 2017 
DistributionIndonesia (Sumatra)

Type locality: Mount Singalang, West Sumatra (elevation 2800 feet) = Gunung Singgalang, Kabupaten Agam, Sumatera Barat, Indonesia, 0.38°S, 100.36°E  
Reproductionoviparous 
TypesLectotype: MSNG 29936, male; paralectotype (?): MRSN (= MSNTO) R2856; MNHN-RA 1889.27 (2133 alpha), also as MHNP, fide Brygoo 1988. 
DiagnosisDiagnosis: The following combination of characters distinguishes Dendragama boulengeri from its congeners: (1) enlarged scales below eye separated from supralabials by continuous row of small lorilabials; (2) enlarged tympanic scale intermediate in size (7–14% of head length) and set far back on head (orbit–meatus 24–32% of head length); (3) sublabial tubercular scales 0–3, usually 1, in front of postrictal modified scale; (4) gulars small, 30–38; (5) dorsal crest of 11–18 projecting scales, serrate with obvious gaps between enlarged projecting scales; (6) dorsolateral crest of more or less continuous posttympanic and dorsolateral series; (7) scales around midbody 62–84; (8) scales of lower flanks heterogenous in size, smooth to feebly keeled; (9) ventrals heavily keeled, 50–63; (10) no white sublabial stripe extending from below eye to level of ear, however 1–3 white spots usually present in this region; (11) most specimens with prominent black prescapular blotch edged posteriorly in white; (12) proximal half of tail with 7–9 dark brown or green bands; (13) buccal epithelium and tongue yellow or orange [Harvey et al. 2017: 83] 
CommentSynonymy partly after WERMUTH 1967. Shaney et al. 2020 formally revalidated D. schneideri from the synonymy of D. boulengeri.

Type species: Calotes boulengeri DORIA 1888 is the type species of the genus Dendragama DORIA 1888.

Key: Shaney et al. 2020 provide a key to the species of Dendragama.

Distribution: see map in Shaney et al 2020: 130 (Fig. 1). 
EtymologyNamed after George Albert Boulenger (1858-1937), herpetologist at the British Museum of Natural History, London. 
References
  • Ahl, E. 1926. Neue Eidechsen und Amphibien. Zool. Anz. 67: 186-192
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Brygoo, E.R. 1988. Les types d’Agamidés (Reptiles, Sauriens) du Muséum national d’Histoire naturelle Catalogue critique. Bull. Mus. Nat. Hist. Nat. 10 (ser. 4) A (3), suppl.: 1-56
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Doria, G. 1888. Note Erpetologiche - Alcuni nuovi Sauri raccolti in Sumatra dal Dr. O. Beccari. Ann. Mus. civ. stor. nat. Genova 2 (ser. 6): 646-652 - get paper here
  • Hallermann, J. 2005. The bizarre arboreal agamids. Reptilia (GB) (42): 8-15 - get paper here
  • Hallermann, J. 2005. Mit Hörnern, Kämmen und Gleithäuten - die bizarren Baumagamen. Reptilia (Münster) 10 (51): 18-25 - get paper here
  • Harvey, Michael B.; Kyle Shaney, Irvan Sidik, Nia Kurniawan, and Eric N. Smith 2017. Endemic Dragons of Sumatra’s Volcanoes: New Species of Dendragama (Squamata: Agamidae) and Status of Salea rosaceum Thominot. Herpetological Monographs 31 (1): 69–97 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Mertens, R. 1954. Über die javanische Eidechse Dendragama fruhstorferi und die Gattung Dendragama. Senckenbergiana 34 (4/6): 185-186
  • Prötzel, D. & J. Forster 2014. Herpetologische Eindrücke von Sumatra. Reptilia (Münster) 19 (109): 80-85
  • Shaney KJ, Harvey MB, Hamidy A, Kurniawan N, Smith EN 2020. Phylogeny and biogeography of Sumatra ́s cloud forest lizards of the genus Dendragama and status of Acanthosaura schneideri. ZooKeys 995: 127–153 - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Uetz, P.; Darko, Y.A.; Voss, O. 2023. Towards digital descriptions of all extant reptile species. Megataxa 010 (1): 027–042 - get paper here
  • Werner, F. 1900. Reptilien und Batrachier aus Sumatra, gesammelt Herrn. Gustav Schneider jr., im Jahre 1897-1898. Zool. Jahrb. (syst) 13: 479-508 - get paper here
 
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