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Diploderma daochengense CAI, ZHANG, LI, DU, XIE, HOU, ZHOU & JIANG, 2022

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Daocheng Mountain Lizard
Chinese: 稻城攀蜥 (dào chéng pān xī) 
SynonymDiploderma daochengense CAI, ZHANG, LI, DU, XIE, HOU, ZHOU & JIANG 2022
Japalura flaviceps — ZHAO et al. 1999: 111 (part.)
Diploderma yulongense — SHU et al. 2021: 259 
DistributionChina (Sichuan)

Type locality: Eya tong Township (28.04754478°N, 100.3224385°E; at elevation 1916 m a.s.l.), Daocheng County, Ganzi Tibetan Autonomous Prefecture, Sichuan Province, China.  
Reproduction 
TypesHolotype: CIB119352, Adult male (filed number DC001), collected by Huaming ZHOU and Jing LI on September 2021.
Allotype: Adult female, CIB119353 (filed number DC003), also collected from Eyatong Ethnic Township (28.0469772°N, 100.32392371°E; at elevation 1930 m a.s.l.), and the same collecting information as the holotype.
Paratypes: DC004 (female, 28.04838729°N, 100.3217799°E; at elevation 1925 m a.s.l.) collected from Eyatong Ethnic Township. The specimens 20210904 (female, 27.946867°N, 100.435083°E; at elevation 1664 m a.s.l.), 20210905 (male, 27.953308°N, 100.427531°E; at elevation 1652 m a.s.l.) collected from Eya Naxi Ethnic Township, Muli Tibetan Autonomous County, Liangshan Yi Autonomous Prefecture, Sichua n Province, China, by Hua ming ZHOU a nd Jing LI on September 2021; 2019ML0037 (male) and 2019ML0038 (female) collected from Xiamaidi Township, Muli Tibetan Autonomous County (27.789794°N, 101.200958°E; at elevation 1889 m a.s.l.) by Feng XIE, Puyang ZHENG and Mingyang CHENG on August 2019; 2019ML0006 (subadult, 28.283388°N, 100.661666°E; a
t elevation 2208 m a.s.l.) and 2019ML0012 (subadult, 28.287653°N, 100.654886°E; at elevation 2158 m a.s.l.) collected from Shuiluo Township, Muli Tibetan Autonomous County by Feng XIE, Puyang ZHENG and Mingyang CHENG on August 2019; ML070706 (female, 28.287653°N, 100.654886°E; at elevation 2158 m a.s.l.) and ML070807 (female, 28.290626°N,100.666664°E; at elevation 2180 m a.s.l.) respectively collected from Shuiluo Township by Feng XIE, Puyang ZHENG and Mingyang CHENG on July 2020. 
DiagnosisDiagnosis: Diploderma daochengense sp. nov. can be distinguished from other Diploderma species by the following characters: 1) TAL/SVL 206.8%–251.6% (average 226.8%) in males, and 169.5%–208.5% (average 189.7%) in females; 2) TRL/ SVL 40.8%–67.3% in males and 45.6%–70.3% in females; 3) SEL/HL 39.6%–71.2% in males and 37.5%–68.8% in females; 4) T4S 14–18 (average 16) and F4S 20–25 (average 23); 5) no bright transverse stripes or bands on dorsal body; 6) ventral scales distinctively keeled; 7) tympanum concealed; 8) transverse gular fold deep, forming a distinct pouch across throat; 9) no radial stripes around eyes;10) inner-lip coloration smoky white, and coloration of tongue and oral cavity light flesh color; 11) ventrolateral body white to grey or light yellow with yellow, cyan or white spots in males; 12) cyan or dark olive green gular spots present in both sexes in life; 13) upper edge of the dorsolateral stripes smooth, and yellow (normally), brown (when dimly lit or stressed) in males, gray while white or light yellow in females in life; 14) fourth toe with claw reaching eye when hindlimbs adpressed forward; 15) well-developed skin folds under nuchal crest present in males only, vertebral crest discontinuous between nuchal and dorsal sections with a distinct gap. (Cai et al. 2022)


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Comment 
EtymologyThe specific epithet of “Daocheng” refers to a county name of Sichuan, and the new species is found on its southeastern edge. 
References
  • Cai B., Zhang M. H., Li J., Du S. M., Xie F., Hou M., Zhou H. M., Jiang J. P. 2022. Three New Species of Diploderma Hallowell, 1861 (Reptilia: Squamata: Agamidae) from the Shaluli Mountains in Western Sichuan, China. Asian Herpetol Res. 13(4): 205–223 - get paper here
 
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