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Dipsadoboa werneri (BOULENGER, 1897)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Werner's Green Tree Snake 
SynonymLeptodira werneri BOULENGER 1897: 281
Dipsas obtusa WERNER 1895 (nec Dipsas obtusa REUSS 1834)
Dipsadomorphus reticulatus WERNER 1909: 55
Crotaphopeltis werneri LOVERIDGE 1933: 32
Dipsadoboa werneri — LAURENT 1951: 210
Crotaphopeltis werneri werneri LOVERIDGE 1959: 39
Dipsadoboa werneri — RASMUSSEN 1986
Dipsadoboa werneri — BROADLEY & HOWELL 1991: 31
Dipsadoboa werneri — WALLACH et al. 2014: 232
Dipsadoboa werneri — SPAWLS et al. 2018: 532 
DistributionTanzania (Usambara Mountains)

Type locality: Usambara Mountains, Tanzania.  
TypesHolotype: BMNH 1946.1.8.32
Holotype: SMNS 3120 [Dipsadomorphus reticulatus WERNER 1909] 
DiagnosisDiagnosis: An East African forest species of Dipsadoboa which is apparently endemic to the Usambara Mountains and which has the following character combination: dorsals in 19-19-13 (rarely 15) rows; ventrals 221-229 and 212-221, males and females, respec- tively; anal entire; subcaudals 102-111 and 98-106, males and females, respectively; 15-16 + II + 1 maxilary teeth; juveniles with a characteristic pattern of cross-bands, which disappear in the adults which have a more or less uniform ground color (yellow- brown to pale grey) with dark-edged scales. 
CommentSynonym after Rasmussen 1986: 53.

Habitat: fully arboreal (Harrington et al. 2018). 
EtymologyNamed after Professor Dr. Franz Josef Maria Werner (1867-1939), an Austrian explorer, zoologist, and herpetologist who worked at Naturhistorisches Museum Wien. 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G. A. 1897. Description of a new snake from Usambara, German East Africa. Ann. Mag. nat. Hist. (6) 19: 281 - get paper here
  • BRANCH, WILLIAM R.; JULIAN BAYLISS, GABRIELA B. BITTENCOURT-SILVA, WERNER CONRADIE, HANLIE M. ENGELBRECHT, SIMON P. LOADER, MICHELE MENEGON, CRISTÓVÃO NANVONAMUQUITXO, KRYSTAL A. TOLLEY 2019. A new species of tree snake (Dipsadoboa, Serpentes: Colubridae) from ‘sky island’ forests in northern Mozambique, with notes on other members of the Dipsadoboa werneri group. Zootaxa 4646 (3): 541–563 - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Lyakurwa, J.V., Howell, K.M., Munishi, L.K., Treydte, A.C. 2019. Uzungwa Scarp Nature Forest Reserve; a unique hotspot for reptiles in Tanzania. Acta Herpetologica 14 (1): 3-14 - get paper here
  • Lyakurwa, John Valentine 2017. The Reptiles of the Uzungwa Scarp Forest Reserve (USFR): An Updated Checklist with Notes on Dagger-Tooth Vine Snake Xyelodontophis uluguruensis. Journal of East African Natural History 106(2):57-65. - get paper here
  • Menegon, Michele; Nike Doggart, Nisha Owen 2008. The Nguru mountains of Tanzania, an outstanding hotspot of herpetofaunal diversity. Acta Herpetologica 3 (2): 107-127
  • Rasmussen,J.B. 1981. The snakes from the rainforest of the Usambara Mountains, Tanzania: a checklist and key. Salamandra 17 (3-4): 173-188 - get paper here
  • Rasmussen,J.B. 1986. On the taxonomic status of Dipsadoboa werneri (BOULENGER), D. shrevei (LOVERIDGE), and Crotaphopeltis hotamboiea kageleri UTHMÖLLER (Boiginae, Serpentes). Amphibia-Reptilia 7: 51-73 - get paper here
  • Rovero, F., Menegon, M., Fjeldså, J., Collett, L., Doggart, N., Leonard, C., Norton, G., Owen, N., Perkin, A., Spitale, D., Ahrends, A., Burgess, N. D. 2014. Targeted vertebrate surveys enhance the faunal importance and improve explanatory models within the Eastern Arc Mountains of Kenya and Tanzania. Diversity and Distributions. doi: 10.1111/ddi.12246 - get paper here
  • Schlüter, A. & Hallermann, J. 1997. The Type Specimens in the Herpetological Collection of the Staatliches Museum für Naturkunde in Stuttgart. Stuttgarter Beitr. Naturk. Ser. A (553): 1-15 - get paper here
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werner,F. 1909. Beschreibung neuer Reptilien aus dem Kgl. Naturalienkabinett in Stuttgart. Jahreshefte Ver. vaterl. Naturk. Württ. 65: 55-63 - get paper here
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