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Dipsas williamsi (CARILLO DE ESPINOZA, 1974)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Williams' Tree Snake 
SynonymSibynomorphus williamsi CARILLO DE ESPINOZA 1974
Sibynomorphus williamsi — CADLE 2007: 248
Sibynomorphus williamsi — WALLACH et al. 2014: 671
Dipsas williamsi — ARTEAGA et al. 2018 
DistributionPeru (Lima)

Type locality: Peru, Lima, Jicamarca  
Reproductionoviparous 
TypesHolotype: MUSM 2170, male, paratypes: MUSM, 6 females 
DiagnosisDiagnosis: Sibynomorphus williamsi is characterized by a high number of ventral scales (173–182 in males, 181–188 in females), usually 6 supralabials with 3–4 bordering the eye, a relatively short tail (21–24% of total length in males, 19–22% in females), and a head distinctly wider than the neck, with the first dorsal crossband usually extending up onto the wide posterior part of the head. The dorsum is strongly crossbanded from head to tail (35–48 bands; posterior bands might be broken into dorsal and lateral blotches, offset to varying degrees, but they retain their boldness). Anterior crossbands can be twice the width of posterior crossbands, but they are always wider than the pale interspaces, which are 1–2 scale rows wide the length of the body.
Sibynomorphus williamsi differs from other species of the genus in western South America as follows (see also previous species accounts). Sibynomorphus vagrans and S. vagus have fewer ventral scales (160 in each case) and different color patterns. Sibynomorphus oligozonatus has a short, blunt snout; a head that is not distinctly wider than the neck; and fewer ventrals in males (145–163 in three specimens). In S. williamsi, the dorsal crossbands (including posterior blotches) are bold the entire length of the body (Figs. 40, 41), whereas in S. oligozonatus, the anterior crossbands are much more well defined than the posterior blotches. Sibynomorphus petersi and S. oneilli differ from S. williamsi in color patterns (see their species accounts) and in having 8 supralabials with 4–5 touching the eye. (Cadle 2007)


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CommentDiet: primarily molluscs (T. de Lema, pers. comm., 16 Oct 2015).

Habitat: fully arboreal (Harrington et al. 2018).

Distribution: see map in Cadle 2007: 253 (Fig. 43). 
EtymologySibynomorphus williamsi was named for Ernest E. Williams (1914-1998), former curator of herpetology and professor of biology at the Museum of Comparative Zoology, Harvard University. 
References
  • Arteaga A, Salazar-Valenzuela D, Mebert K, Peñafiel N, Aguiar G, Sánchez-Nivicela JC, Pyron RA, Colston TJ, Cisneros-Heredia DF, Yánez-Muñoz MH, Venegas PJ, Guayasamin JM, Torres-Carvajal O 2018. Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79-147 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Cadle, J.E. 2007. The snake genus Sibynomorphus (Colubridae: Dipsadinae: Dipsadini) in Peru and Ecuador, with comments on the systematics of Dipsadini. Bull. Mus. Comp. Zool. Harvard 158(5):183-283 - get paper here
  • Carillo DE ESPINOZA 1974. Sibynomorphus williamsi nov. sp. (Serpentes: Colubridae. Publicaciones del Museo de Historia Natural ‘‘Javier Prado,’’ serie A (Zoología), 24: 1-16
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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