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Draco maculatus (GRAY, 1845)

IUCN Red List - Draco maculatus - Least Concern, LC
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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
SubspeciesDraco maculatus divergens TAYLOR 1934
Draco maculatus whiteheadi BOULENGER 1900
Draco maculatus maculatus (GRAY 1845)
Draco maculatus haasei BOETTGER 1893 
Common NamesE: Spotted Flying Dragon, Spotted Gliding Lizard
G: Gefleckter Flugdrache
Chinese: 斑飞蜥 
SynonymDracunculus maculatus GRAY 1845
Draco maculatus — CANTOR 1847
Dracunculus maculatus — GÜNTHER 1861: 268
Draco maculatus — BOULENGER 1885: 262
Draco haasei BOETTGER 1893: 429 (fide SMITH 1935)
Draco maculatus — BOULENGER 1885: 262
Draco maculatus — WERNER 1910: 10
Draco maculatus — SMITH 1935: 138
Draco maculatus — TAYLOR 1963: 823
Draco maculatus — WERMUTH 1967
Draco maculatus — MANTHEY & GROSSMANN 1997: 169
Draco maculatus — COX et al. 1998: 100
Draco maculatus — MCGUIRE & KIEW 2001
Draco cf. maculatus — ZIEGLER et al. 2015
Draco maculatus — WANG et al. 2022

Draco maculatus divergens TAYLOR 1934
Draco divergens TAYLOR 1934
Draco maculatus divergens — TAYLOR 1963
Draco maculatus divergens — WERMUTH 1967
Draco maculatus divergens — MUSTERS 1983
Draco maculatus divergens — NGUYEN et al. 2009
Draco maculatus divergens — KLABACKA et al. 2020

Draco maculatus haasei BOETTGER 1893
Draco haasei BOETTGER 1893: 429
Draco hasei — BOULENGER 1894 (Zool. Rec., in error)
Draco maculatus haasii — SMITH & KLOSS 1915 (in error)
Draco maculatus haasei — TAYLOR 1963: 829
Draco maculatus haasei — WERMUTH 1967: 51
Draco maculatus haasei — MUSTERS 1983
Draco maculatus haasii — HALLERMANN 2005
Draco maculatus haasei — NGUYEN et al. 2009
Draco maculatus haasei — KLABACKA et al. 2020

Draco maculatus maculatus (GRAY 1845)
Draco maculatus maculatus — HENNIG 1936
Draco maculatus maculatus — TAYLOR 1963
Draco maculatus maculatus — WERMUTH 1967: 50
Draco maculatus maculatus — MUSTERS 1983
Draco maculatus maculatus — NGUYEN et al. 2009
Draco maculatus maculatus — KLABACKA et al. 2020

Draco maculatus whiteheadi BOULENGER 1900
Draco whiteheadi BOULENGER 1899: 956
Draco whiteheadi — BARBOUR 1909: 63
Draco whiteheadi — SMTH 1923
Draco maculatus whiteheadi — HENNIG 1936
Draco maculatus whiteheadi — TAYLOR 1963: 831
Draco whiteheadi — COLBERT 1967
Draco maculatus whiteheadi — MUSTERS 1983
Draco maculatus whiteheadi — NGUYEN et al. 2009
Draco maculatus whiteheadi — KLABACKA et al. 2020 
DistributionS China (Fujian, Hainan, Guangxi, Yunnan, SE Xizang = Tibet),
India (E. Himalayas to Assam - Naga hills and Abor country, Mizoram), Myanmar (= Burma), Laos, Vietnam (Son La etc.), Thailand, W Malaysia (incl. Pulau Langkawi)

divergens: NW Thailand (Chiang Mai Valley); Type locality: Chieng Mai, N Siam; restricted to Doi Suthep Mountain by TAYLOR 1963. Range restricted to Chiang Mai Valley by Klabacka et al. 2020.

haasei: E Thailand, Cambodia, S Vietnam; Type locality: Chantaboon, Siam. May be restricted to Hainan fide Klabacka et al. 2020.

whiteheadi: N Vietnam, Hainan; Type locality: Five-finger Mountains, interior of Hainan.

Type locality: Penang  
Reproductionoviparous 
TypesHolotype: BMNH 1946.8.27.22 (23.9a) fide Musters 1983: 53
Holotype: ANSP 19938; Thailand, Chiang Mai [divergens]
Holotype: SMF 9700, male [haasei]
Holotype: BMNH 1946.8.26.79 [whiteheadi] 
DiagnosisDiagnosis: A moderately sized species (maximum snout-vent length 87 mm); nostril directed outward and slightly upward; a series of keeled scales for- ming a A-shaped figure on the snout; usually two median teeth in the upper jaw; supralabials 7-11, smooth; a low thornlike scale on the supraciliary edge; tym- panum covered with small scales; gular pouch in males with slightly enlarged scales; lateral pouches with enlarged scales; nuchal crest present; dorsals 117-152, some are keeled, unequal; usually five ribs in patagium; hindlimbs about as long as distance between the legs; scales on tail strongly keeled, more or less with a caudal crest in males (Musters 1983: 52).


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CommentDistribution: possibly in Bhutan (Lenz 2012). ZHAO AND ADLER (1993) question the validity of the type locality.

Phylogenetics: although KLABACKA et al. 2020 presented a phylogenetic analysis of the various population of Draco maculatus, they refrained from elevating any of the subpspecies or changing their status. 
References
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