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Enyalioides cofanorum DUELLMAN, 1973

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Higher TaxaHoplocercidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Cofan woodlizard, Duellman's Dwarf Iguana
S: lagartijas de palo cofanes 
SynonymEnyalioides cofanorum DUELLMAN 1973
Enyalioides cofanorum — DUELLMAN 1978: 203
Enyalioides cofanorum — LEWIS 2002
Enyalioides cofanorum — TORRES-CARVAJAL et al. 2011 
DistributionColombia [Castro,F. (pers. comm.)], Ecuador, N Peru ?

Type locality: Ecuador, Napo, Santa Caecilia [0°3'N, 76°59'33''W]  
Reproductionoviparous 
TypesHolotype: KU 146658 
DiagnosisDiagnosis: This species can be distinguished from other species of Enyalioides by the combination of the following characters: scattered, projecting, large dorsal scales (inconspicuous or absent in some juvenile and male individuals) between fore and hind limbs; strongly keeled ventrals; large dark patch on posterior aspect of gular region in both sexes; and light stripe extending from postympanic region to scapular region. The only other species of Enyalioides with scattered, projecting dorsal scales is E. heterolepis, from which E. cofanorum differs in lacking projecting scales on the hind limbs. In addition, males and females of E. cofanorum (maximum SVL = 107 mm and 109 mm, respectively) are smaller than those of E. heterolepis (maximum SVL = 137 mm and 115 mm, respectively), and these two species occur on opposite sides of the Andes. Among other species occuring east of the Andes, specimens of E. cofanorum lacking the projecting dorsal scales are similar morphologically to specimens of E. microlepis (character states in parentheses) and can be distinguished from them by having conspicuous dorsolateral crests between hind limbs (inconspicuous or absent), and a smaller body size (maximum SVL = 127 mm and 116 mm in males and females, respectively) [from TORRES-CARVAJAL et al. 2011]. 
Comment 
EtymologyNamed after the Cofan Indians. Not in Peru fide TORRES-CARVAJAL et al. (2008) 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Dixon, J. R.; Soini, P. 1975. The reptiles of the upper Amazon basin, Iquitos region, Peru, Part I. Lizards and Amphisbaenians. Milwaukee Public Museum Contributions in Biology and Geology 4: 1-58 [1986?] - get paper here
  • Duellman, W. E. 1978. The biology of an equatorial herpetofauna in Amazonian Ecuador. Misc. Publ. Univ. Kans. Mus. Nat. Hist. 65: 1-352 - get paper here
  • Duellman, W.E. 1973. Descriptions of new lizards from the upper Amazon Basin. Herpetologica 29: 228-231 - get paper here
  • Duellman,W.E. & Mendelson III, J.R. 1995. Amphibians and reptiles from Northern Departamento Loreto, Peru: Taxonomy and Biogeography. Univ. Kansas Sci. Bull. 55 (10): 329-376 - get paper here
  • Lewis, R.R. 2002. Threats facing endemic herpetofauna in the cloud forest reserves of Ecuador. Herpetological Bulletin (79): 18-26 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Torres-Carvajal, O.; Almenáriz, A.; Valencia, J.; Yúnez-Muñoz, M. & Reyes, J.P. 2008. A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Pap. Avulsos Zool. (São Paulo) 48 (20): 227-235 - get paper here
  • Torres-Carvajal, Omar; Kevin de Queiroz & Richard Etheridge 2009. A new species of iguanid lizard (Hoplocercinae, Enyalioides) from southern Ecuador with a key to eastern Ecuadorian Enyalioides. ZooKeys 27: 59–71 - get paper here
  • Torres-Carvajal, Omar; RICHARD ETHERIDGE & KEVIN DE QUEIROZ 2011. A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania). Zootaxa 2752: 1–44 - get paper here
  • Venegas PJ, Torres-Carvajal O, Duran V, de Queiroz K 2013. Two sympatric new species of woodlizards (Hoplocercinae, Enyalioides) from Cordillera Azul National Park in northeastern Peru. ZooKeys 277: 69–90. doi: 10.3897/zookeys.277.3594 - get paper here
  • Wiens, John J. and Richard E. Etheridge 2003. Phylogenetic relationships of Hoplocercid lizards: coding and combining meristic, morphometric, and polymorphic data using step matrices. Herpetologica 59 (3): 375-398 - get paper here
 
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