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Enyalioides palpebralis (BOULENGER, 1883)

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Higher TaxaHoplocercidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: horned woodlizard, Boulenger's Dwarf Iguana
S: lagartijas de palo cornudas 
SynonymEnyalius palpebralis BOULENGER 1883: 46
Enyalioides palpebralis — BOULENGER 1885: 116
Enyalioides palpebralis — PETERS & DONOSO-BARROS 1970: 115
Enyalioides palpebralis — DIRKSEN & DE LA RIVA 1999
Enyalioides palpebralis — TORRES-CARVAJAL et al. 2011 
DistributionE Peru, Brazil (Acre), N Bolivia

Type locality: Cashiboya, E Peru [ 7°32'60''S, 74°52'60''W], 160 m elevation  
Reproductionoviparous 
TypesHolotype: BMNH 81.5.13.25 
DiagnosisDiagnosis: This species is different from other species of Enyalioides in having a superciliary triangular flap projecting posterolaterally over each eye; a wide postorbital process of the squamosal (process narrow in other species); a pair of enlarged dorsal tubercles on the posterolateral aspect of the parietal roof; and in lacking a posteriorly projecting squamosal process of the postorbital (Fig. 8; Wiens & Etheridge 2003). In addition most specimens of E. palpebralis have a discontinuous vertebral crest, with a small gap on the neck, and most lack femoral pores [from TORRES-CARVAJAL et al. 2011]. 
Comment 
EtymologyNamed after Latin palpebralis, of or on the eyelids. (BOULENGER 1883, Esteban Lavilla, pers. comm., April 2024) 
References
  • Avila-Pires, T.C.S. 1995. Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen 299: 1-706 - get paper here
  • Bernarde, Paulo Sérgio;, Reginaldo Assêncio Machado & Luiz Carlos Batista Turci 2011. Herpetofauna da área do Igarapé Esperança na Reserva Extrativista Riozinho da Liberdade, Acre – Brasil. Biota Neotropica 11 (3): 117-144 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Boulenger, George A. 1883. Description of a new species of lizard of the genus Enyalius. Proc. Zool. Soc. London 1883: 46 - get paper here
  • CATENAZZI, A., LEHR, E. & VON MAY, R. 2013. The amphibians and reptiles of Manu National Park and its buffer zone, Amazon basin and eastern slopes of the Andes, Peru. Biota Neotropica 13 (4): 269-283
  • Crnobrna B, Santa-Cruz Farfan R, Gallegos C, López-Rojas JJ, Llanqui IB, Pisco GP, Arbaiza AK 2023. Herpetological records from the Abujao basin, central Peruvian Amazon. Check List 19(4): 433-465 - get paper here
  • Dirksen, L. & De la Riva, I. 1999. The lizards and amphisbaenians of Bolivia (Reptilia, Squamata): checklist, localities, and bibliography. Graellsia 55: 199-215 - get paper here
  • Duellman, W. E. 2005. Cusco Amazónico: The Lives of Amphibians and Reptiles in an Amazonian Rainforest. Comstock Pub Assoc.
  • Fonseca, Wirven Lima da; Julivaldo Dantas da Silva, Arthur Diesel Abegg, Conrado Mario da Rosa, Paulo Sergio Bernarde 2019. Herpetofauna of Porto Walter and surrounding areas, Southwest Amazonia, Brazil. Herpetology Notes 12: 91-107 - get paper here
  • Freitas, M. A. de, Venancio, N. M., Abegg, A. D., Azevedo, W. S, Pereira, V. O., Zanotti, A. P., Veloso, A., Schwarzbach, L., Oliveira, A. G., da Silva, R. C. C., de Amorim, V. R. G. and de Moura, G. J. B. 2020. Herpetofauna at the Rio Acre Ecological Station, Amazon Rainforest, Brazil. Herpetology Notes 13: 33-48. - get paper here
  • Moravec J., Aparicio J. 2005. Notes on the herpetofauna of Bioceanica and Bolpebra (Provincia Nicolas Suaréz, Departamento Pando, Biolivia). Journal of the National Museum (Prague), Natural History Series 174 (1-4): 95-113
  • Peters, James A. & Donoso-Barros, Roberto 1970. Catalogue of the Neotropical Squamata: Part II. Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 293 pp. - get paper here
  • Pianka, E.R. & Vitt, L.J. 2003. Lizards - Windows to the Evolution of Diversity. University of California Press, Berkeley, 347 pp. [review in Copeia 2004: 955] - get paper here
  • Prudente, A.L.C.; F. Magalhães; A. Menks; J.F.M. Sarmento. 2013. Checklist of Lizards of the Juruti, state of Pará, Brazil. Check List 9 (1):42-50 - get paper here
  • Rabosky, Daniel L.; Rudolf von May, Michael C. Grundler and Alison R. Davis Rabosky 2019. The Western Amazonian Richness Gradient for Squamate Reptiles: Are There Really Fewer Snakes and Lizards in Southwestern Amazonian Lowlands? Diversity 11: 199; doi:10.3390/d11100199 - get paper here
  • RIBEIRO-JÚNIOR, MARCO A. 2015. Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. I. Dactyloidae, Hoplocercidae, Iguanidae, Leiosauridae, Polychrotidae, Tropiduridae. Zootaxa 3983 (1): 001–110 - get paper here
  • Ribeiro-Júnior, Marco A. & Silvana Amaral 2016. Diversity, distribution, and conservation of lizards (Reptilia: Squamata) in the Brazilian Amazonia. Neotropical Biodiversity, 2:1, 195-421 - get paper here
  • Salas, Antonio W. 1995. HERPETOFAUNA PERUANA: UNA VISION PANORAMICA SOBRE INVESTIGACION, CONSERVACION Y MANEJO. Biotempo 2: 125 - get paper here
  • Torres-Carvajal, Omar; RICHARD ETHERIDGE & KEVIN DE QUEIROZ 2011. A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania). Zootaxa 2752: 1–44 - get paper here
  • Venegas PJ, Torres-Carvajal O, Duran V, de Queiroz K 2013. Two sympatric new species of woodlizards (Hoplocercinae, Enyalioides) from Cordillera Azul National Park in northeastern Peru. ZooKeys 277: 69–90. doi: 10.3897/zookeys.277.3594 - get paper here
  • Wiens, John J. and Richard E. Etheridge 2003. Phylogenetic relationships of Hoplocercid lizards: coding and combining meristic, morphometric, and polymorphic data using step matrices. Herpetologica 59 (3): 375-398 - get paper here
 
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