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Geckolepis typica GRANDIDIER, 1867

IUCN Red List - Geckolepis typica - Least Concern, LC
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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Grandidier's Gecko
G: Schuppengecko 
SynonymGeckolepis typicus GRANDIDIER 1867: 233
Geckolepis typica — BOULENGER 1885: 192
Geckolepis typica var. anomala MOCQUARD 1909
Geckolepis anomala — MOCQUARD 1909: 18
Geckolepis typica var. modesta METHUEN & HEWITT 1913: 186
Geckolepis typica modesta — ANGEL 1942
Geckolepis typica — ANGEL 1942: 43
Geckolepis anomala — ANGEL 1942: 42
Geckolepis petiti ANGEL 1942: 42
Geckolepis petiti — WERMUTH 1965: 31
Geckolepis anomala — WERMUTH 1965: 30
Geckolepis modesta — KLUGE 1991: 8
Geckolepis anomala — KLUGE 1993
Geckolepis petiti — KLUGE 1993
Geckolepis petiti — GLAW & VENCES 1994: 275
Geckolepis typica — GLAW & VENCES 1994: 275
Geckolepis anomala — GLAW & VENCES 1994: 275
Geckolepis typica modesta — GLAW & VENCES 1994: 275
Geckolepis petiti — RÖSLER 2000: 79
Geckolepis typica modesta — RÖSLER 2000: 80
Geckolepis anomala — RÖSLER 2000: 79
Geckolepis typica — RÖSLER 2000: 79
Geckolepis cf. typica — D’CRUZE et al. 2009
Geckolepis cf. petiti — D’CRUZE et al. 2009
Geckolepis typica — HAWLITSCHEK et al. 2015 
DistributionMadagascar, Saint Marie, Ile aux Prunes

Type locality: Sainte Marie, Madagascar.

anomala: Type locality: Androy Nord, Madagascar.

petiti: Type locality: Andranovaho, Mahafaly Province, Madagascar.  
Reproductionoviparous 
TypesLectotype: MNHN-RA 1994.1748 (former MNHN-RA 1454A) (designation by KÖHLER et al. 2009); paralectotypes: MNHN-RA 1454, MNHN-RA 8119, and MNHN-RA 1994.1749 (former MNHN-RA 1454B).
Types: DNMNH (= TM) 4054–5, 4146 [modesta] 
DiagnosisDiagnosis (genus): A genus of the family Gekkonidae that is easily recognized by the large flat imbricate cycloid (fish-scale) body scales that are shed easily—the cycloid scales and most of the underlying dermis can be autotomized; vertical pupil; digits strongly dilated with transverse undivided subdigital lamellae; all digits with a retractile claw; no preanal or femoral pores in males. [from KÖHLER et al. 2009].


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CommentSynonymy: KÖHLER et al. 2009 synonymized anomala and modesta with typica.

Type species: Geckolepis typicus GRANDIDIER 1867 is the type species of the genus Geckolepis GRANDIDIER 1867.

Diversity: LEMME et al. (2013) described other Confirmed Candidate Species, based on genetic differences, which exhibit only little morphological diversity.

Behavior: nocturnal

Reference images: see Uetz et al. 2024 for high-resolution reference images for this species. 
EtymologyNamed after the Latin adjective typicus, supposedly referring to the species' morphology which was considered as being typical for Geckolepis.

The genus was named after gecko and Greek lepis = scale, for the unusual scalation in this genus.  
References
  • Angel, F. 1942. Les Lézards de Madagascar. Mem. Acad. Malagache, Tananarive XXXVI: 193 pp.
  • Bauer, A.M.; Russell, A.P.; Shadwick, R.E. 1993. Skin mechanics and morphology of 2 species of Pachydactylus (Reptilia, gekkonidae). South African J Zoology 28 (4): 192-197 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Cocca, W., G. M. Rosa, F. Andreone, G. Aprea, P. E. Bergò, F. Mattioli, V. Mercurio, J. E. Randrianirina, D. Rosado, M. Vences & A. Crottini 2018. The herpetofauna (Amphibia, Crocodylia, Squamata, Testudines) of the Isalo Massif, Southwest Madagascar: combining morphological, molecular and museum data. Salamandra 54 (3): 178-200 plus Supplementary Material - get paper here
  • D'Cruze, Neil; Annette Olsonn, David Henson, Sunil Kumar, and David Emmett. 2009. The Amphibians and Reptiles of the Lower Onilahy River Valley, a Temporary Protected Area in Southwest Madagascar. Herp. Cons. Biol. 4: 62-79 - get paper here
  • Evans, A. M., Marshal, J. P., & Alexander, G. J. 2021. Forest patch characteristics affect reptile occurrence in north‐western Madagascar. Austral Ecology - get paper here
  • Gardner, Charlie J.; Xavier Vincke, Louise D. Jasper 2015. Additions to the urban herpetofauna of Toliara, southwest Madagascar. Herpetology Notes 8: 47-49 - get paper here
  • Glaw ,F. & Vences, M. 1994. A Fieldguide to the Amphibians and Reptiles of Madagascar. Vences & Glaw Verlag, Köln (ISBN 3-929449-01-3)
  • Grandidier, A. 1867. Liste des reptiles nouveaux découverts, en 1866, sur la côte sud-ouest de Madagascar. Revue et Magazine de Zoologie (Paris), Sér. 2, 19: 232-234
  • Hawlitschek, Oliver; Mark D. Scherz; Nicolas Straube, Frank Glaw 2015. Resurrection of the Comoran fish scale gecko Geckolepis humbloti Vaillant, 1887 reveals a disjunct distribution caused by natural overseas dispersal. Organisms Diversity & Evolution, doi:10. 1007/ s13127-015-0255-1 - get paper here
  • Köhler, Gunther; Hans-Helmut Diethert, Ronald A. Nussbaum, and Christopher J. Raxworthy 2009. A REVISION OF THE FISH SCALE GECKOS, GENUS GECKOLEPIS GRANDIDIER (SQUAMATA, GEKKONIDAE) FROM MADAGASCAR AND THE COMOROS. Herpetologica 65 (4): 419-435 - get paper here
  • Lemme, Inga; Martina Erbacher, Nathalie Kaffenberger, Miguel Vences & Jörn Köhler 2013. Molecules and morphology suggest cryptic species diversity and an overall complex taxonomy of fish scale geckos, genus Geckolepis. Org Divers Evol 13:87-95 - get paper here
  • Mashinini, P. L. and Mahlangu, L. M. 2013. An annotated catalogue of the types of gekkonid lizards (Reptilia: Squamata: Gekkonidae) in the Herpetology collection of the Ditsong National Museum of Natural History, South Africa. Annals of the Ditsong National Museum of Natural History 3: 165-181
  • Meier, H. 1980. Geckolepis, eine besondere Gecko-Gattung Madagaskars. Herpetofauna 2 (5): 11-14 - get paper here
  • Methuen, P. A. & J. Hewitt 1913. On a collection of reptiles from Madagascar made during the year 1911. Annals Transvaal Mus. 3 (4): 183-193. - get paper here
  • Mezzasalma, M., F. Andreone, F. Glaw, G. Odierna, A. Petraccioli & F. M. Guarino 2018. Chromosome aneupolyploidy in an endemic Malagasy gecko (Gekkonidae: Geckolepis). Salamandra 54 (1): 56-62 - get paper here
  • Mocquard, F. 1902. Sur une collection de reptiles et de batraciens recueillis par M. Alluaud dans le sud de Madagascar. Bulletin de la Sociéte Philomathique de Paris, 4:5—21 - get paper here
  • Mocquard, François 1909. Synopsis des familles, genres et espèces des reptiles écailleux et des batraciens de Madagascar. Nouvelles Archives du Museum National d'Histoire Naturelle, Paris (Paris), 5e Ser., 1: 1-108. - get paper here
  • Radonirina, Oninjatovo; H.; Randriamahatantsoa, B.; Rabibisoa, N.H.C.; Raxworthy, C.J. 2023. Amphibians and Reptiles of the Montagne des Français: An Update of the Distribution and Regional Endemicity. Animals 13: 3361 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Uetz, P.H.; Patel, M.; Gbadamosi, Z.; Nguyen, A.; Shoope, S. 2024. A Reference Database of Reptile Images. Taxonomy 4: 723–732 - get paper here
 
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