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Gekko intermedium (TAYLOR, 1915)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Intermediate Flying Gecko 
SynonymPtychozoon intermedium TAYLOR 1915: 94
Ptychozoon intermedia — TAYLOR 1922
Ptychozoon intermedium — KLUGE 1993
Ptychozoon intermedium — RÖSLER 2000: 107
Gekko (Ptychozoon) intermedium — WOOD et al. 2019
Gekko (Ptychozoon) intermedium — WOOD et al. 2020 
DistributionPhilippines (Mindanao, Leyte, Dinagat, Maripipi)

Type locality: Bunauan, Agusan Province, Mindanao.
Type locality: Malagos public water reservoir compound and Philippine Eagle Research and Nature Center, Barangay Malagos, Baguio district, Municipality of Calinan, Cavao City Province, Mindanao [neotype].  
Reproductionoviparous 
TypesNeotype: PNM 2501 (designated by Brown et al. 1997); original type: Bureau of Science collection No. 1776 from Bunauan, Agusan Province, Mindanao, July 12, 1912; E. H. Taylor, collector. 
DiagnosisDiagnosis: “Large scales of the dermal lobe on the side of the head equal to, or slightly smaller than, the largest abdominal scales; back with from 10 to 12 irregular longitudinal rows of flat round tubercles; tail with a narrow unsegmented terminal flap, no wider than the nearest lobes, slightly notched behind; annulations on the tail marked by lateral lobes directed somewhat backward, and a series of 4 or more enlarged scales above, much larger than the scales on the lateral lobes.” (Taylor 1915: 94)


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Comment 
EtymologyNamed for being intermediate between (Ptychozoon) kuhli and P. horsfieldii (spelled as “horsfeldii” by Taylor 1915). 
References
  • Beukema, W. 2011. Herpetofauna of disturbed forest fragments on the lower Mt. Kitanglad Rnage, Mindanao Isand, Philippines. Salamandra 47 (2): 90-98 - get paper here
  • Brown, Rafe M. 1999. New species of parachute gecko (Squamata: Gekkonidae: genus Ptychozoon) from northeastern Thailand and central Vietnam. Copeia 1999 (4): 990-1001 - get paper here
  • Brown, Rafe M., John W. Ferner and Arvin C. Diesmos. 1997. Definition of the Philippine Parachute Gecko, Ptychozoon intermedium Taylor 1915 (Reptilia: Squamata: gekkonidae): Redescription, designation of a neotype, and comparisons with related species. Herpetologica 53 (3): 357-373. - get paper here
  • Brown, Rafe M.; Cameron D. Siler, L. Lee Grismer, Indraneil Das, Jimmy A. McGuire 2013. Phylogeny and cryptic diversification in Southeast Asian flying geckos. Molecular Phylogenetics and Evolution 66 (2):351-361 [2012] - get paper here
  • Das, I. & Vijayakumar, S.P. 2009. New species of Ptychozoon (Sauria: Gekkonidae) from the Nicobar Archipelago, Indian Ocean. Zootaxa 2095: 8–20 - get paper here
  • GRISMER, L. LEE; PERRY L. JR. WOOD, MYINT KYAW THURA, MARTA S. GRISMER, RAFE M. BROWN, BRYAN L. STUART 2018. Geographically structured genetic variation in Ptychozoon lionotum (Squamata: Gekkonidae) and a new species from an isolated volcano in Myanmar. Zootaxa 4514 (2): 202–214 - get paper here
  • Manthey, U. 1982. Die Gattung Ptychozoon, Teil 2 - Mit einem Bestimmungsschlüssel für die fünf Arten. Sauria 4 (3): 5-12 - get paper here
  • Manthey, U. 1982. Die Gattung Ptychozoon, Teil 1 - Mit einem Bestimmungsschlüssel für die fünf Arten. Sauria 4 (2): 11-17 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Sanguila MB, Cobb KA, Siler CD, Diesmos AC, Alcala AC, Brown RM 2016. The amphibians and reptiles of Mindanao Island, southern Philippines, II: the herpetofauna of northeast Mindanao and adjacent islands. ZooKeys 624: 1–132, doi: 10.3897/zookeys.624.9814 - get paper here
  • SUMONTHA, MONTRI; OLIVIER S.G. PAUWELS, KIRATI KUNYA, CHAIWAT LIMLIKHITAKSORN, SIRICHAI RUKSUE, APIRAT TAOKRATOK, MICHEL ANSERMET & LAWAN CHANHOME 2012. A new species of Parachute Gecko (Squamata: Gekkonidae: genus Ptychozoon) from Kaeng Krachan National Park, western Thailand. Zootaxa 3513: 68–78 - get paper here
  • Sy, Emerson Y., Con Foley and Rommel M. Cruz. 2014. Ptychozoon intermedium (Philippine flying gecko) avian predation. Herpetological Review 45 (4): 698-699 - get paper here
  • Taylor, E.H. 1915. New species of Philippine lizards. Philippine Journal of Science. 10: 89-109 - get paper here
  • Taylor, E.H. 1922. The lizards of the Philippine Islands. Department of Agriculture and Natural Resources, Bureau of Science, Government of the Philippine Islands, Manila, Publication no. 17: 269 pp. - get paper here
  • Wood Jr, Perry L.; Xianguang Guo, Scott L. Travers, Yong-Chao Su, Karen V. Olson, Aaron M. Bauer, L. Lee Grismer, Cameron D. Siler, Robert G. Moyle, Michael J. Andersen, Rafe M. Brown 2019. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. bioRxiv 717520 [preprint] - get paper here
  • Wood, P. L., Guo, X., Travers, S. L., Su, Y. C., Olson, K. V., Bauer, A. M., Grismer, L. L., Siler, C. D., Moyle, R. G., Andersen, M. J. and Brown, R. M. 2020. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Molecular Phylogenetics and Evolution 146: 106731 [corrigendum: MPE 164: 107255] - get paper here
 
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