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Gekko nadenensis LUU, NGUYEN, LE, BONKOWSKI & ZIEGLER, 2017

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Naden Gecko
Laotian: Kap Ke Naden
G: Naden Gecko 
SynonymGekko nadenensis LUU, NGUYEN, LE, BONKOWSKI & ZIEGLER 2017
Gekko (Japonigekko) nadenensis — WOOD et al. 2019
Gekko (Japonigekko) nadenensis — WOOD et al. 2020 
DistributionC Laos (Khammouane Province)

Type locality: on a karst cliff, near the entrance of Nang Log cave (17°30’N, 105°23’E, elevation 178 m a.s.l.), Naden Village, Gnommalath District, Khammouane Province, central Laos  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: VNUF R.2016.1, adult male, collected by Vinh Quang Luu and Kieusomphone Thanabuaosy on 20 February 2016. Paratype. NUOL-R.2016.2, adult male, the same data as the holotype; ZFMK 98741, gravid adult female, collected on a karst cliff inside Nang Log cave by Vinh Quang Luu, Thomas Calame, and Kieusomphone Thanabuaosy on 29 March 2015. 
DiagnosisDiagnosis: Gekko nadenensis sp. nov. differs from its relatives by a combination of the following characters: a medium-sized gecko species (SVL 61.0–77.1 mm); nares bordered with rostral; internasals absent; postmentals enlarged; interorbital scales between anterior corners of the eyes 28–30; dorsal tubercles absent; ventral scales between mental and cloacal slit 175–185; midbody scale rows 123–140; ventral scale rows 38–40; subdigital lamellae on first toes 13–15, on fourth toes 14–16; finger and toe webbing present at base; tubercles on upper surface of fore and hind limbs absent; precloacal pores six (3+3 or 5+1) in a discontinuous row in males and absent in the female; postcloacal tubercles 1 or 2; tubercles absent on dorsal surface of tail base; subcaudals distinctly enlarged; dorsal surface of body with greyish brown blotches.


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CommentSimilar species: Molecular analyses demonstrated the new species is closely related to G. bonkowskii and G. thakhekensis, but separated from them by approximately 7% in genetic divergence as shown by a fragment of the mitochondrial ND2 gene

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyNamed after its type locality, Naden Village 
References
  • LUU, VINH QUANG; TRUONG QUANG NGUYEN, MINH DUC LE, MICHAEL BONKOWSKI, THOMAS ZIEGLER 2017. A new karst dwelling species of the Gekko japonicus group (Squamata: Gekkonidae) from central Laos. Zootaxa 4263 (1): 179-193 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Wood Jr, Perry L.; Xianguang Guo, Scott L. Travers, Yong-Chao Su, Karen V. Olson, Aaron M. Bauer, L. Lee Grismer, Cameron D. Siler, Robert G. Moyle, Michael J. Andersen, Rafe M. Brown 2019. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. bioRxiv 717520 [preprint] - get paper here
  • Wood, P. L., Guo, X., Travers, S. L., Su, Y. C., Olson, K. V., Bauer, A. M., Grismer, L. L., Siler, C. D., Moyle, R. G., Andersen, M. J. and Brown, R. M. 2020. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Molecular Phylogenetics and Evolution 146: 106731 [corrigendum: MPE 164: 107255] - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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