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Gekko tokehos (GRISMER, WOOD, GRISMER, QUAH, THY, PHIMMACHAK, SIVONGXAY, SEATEUN, STUART, SILER, MULCAHY, ANAMZA & BROWN, 2019)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Cambodian Parachute Gecko 
SynonymPtychozoon tokehos GRISMER, WOOD, GRISMER, QUAH, THY, PHIMMACHAK, SIVONGXAY, SEATEUN, STUART, SILER, MULCAHY, ANAMZA & BROWN 2019: 179
Gekko (Ptychozoon) tokehos — WOOD et al. 2019
Gekko (Ptychozoon) tokehos — WOOD et al. 2020 
DistributionCambodia (Kampong Speu), Thailand, S Vietnam

Type locality: Kirirom National Park, Phnom Sruoch District, Kampong Speu Province, Cambodia (11.32611°N, 104.06556°E, 700 m in elevation).  
Reproduction 
TypesHolotype. FMNH 261853 adult female, collected by Bryan L. Stuart and Dara Anon 5 June 2000. Paratypes. FMNH 261851–52 bear the same collection data as the holotype except they were collected on 3 June 2000. FMNH 261854 bears the same collection data as the holotype except it was collected on 5 November 2000 by Joe Walston. CBC 03162 adult female collected by Neang Thy on 9 June 2017 from Bokor National Park, Teuk Chou District, Kampot Province, Cambodia (10.60156°N, 104.03200°E, 325 m in elevation). NCSM 98986 adult female collected by Neang Thy on 10 June 2017 from Bokor National Park, Teuk Chou District, Kampot Prov- ince, Cambodia (10.60372°N, 104.02944°E, 328 m in elevation). NCSM 98987 adult female collected by Neang Thy on 12 June 2017 from Bokor National Park, Teuk Chou District, Kampot Province, Cambodia (10.58753°N, 104.03269°E, 315 m in elevation). FMNH 177359 adult male collected by Edward H. Taylor on 11 June 1969 from the Khao Chong Reserve, Trang Province, Thailand (7.54350°N 99.79800°E, 127 m in elevation). FMNH 181844 adult male collected by W. Ronald Heyer on 11 June 1969 from Pak Thong Chai District, Nakhon Ratchasima Prov- ince, Sakaerat (Environmental Research Station), Thailand (14.50000°N, 100.86670°E, 16 m in elevation). 
DiagnosisDiagnosis: Ptychozoon tokehos sp. nov. differs from all other species of Ptychozoon by having the following unique combination of characters: a maximum SVL of 97.5 mm; supranasals not in contact; 8–11 supralabials; 10–12 infralabials; infra-auricular cutaneous flap; weak supra-auricular ridge present; no dorsal or caudal tubercles; imbricate parachute support scales on dorsal surface of patagia; prominently raised ridges on ventral surface of patagia; 80–95 midbody dorsal scales; 30–37 ventral scales; an emargination between the pre-antebrachial flap and digit I; no enlarged femoral scales; 20–24 pore-bearing precloacal scales in males; 18–25 enlarged precloacal scales; 4–7 rows of enlarged post-precloacal scales; 13–18 transverse subdigital lamellae on the fourth toe; approximately 28–34 scales across the widest portion of the caudal flap; enlarged dorsal caudal scales forming intermittent whorls; distal lobes fusing to form a short, narrow, caudal flap; edges of caudal flap smooth; caudal lobes angled posteri- orly; caudal lobes variably showing posterior reduction in size; postorbital striping variable; four dark body bands between limb insertions; and irregularly shaped, white, vertebral markings usually not present (absent in 14 of 16 specimens; Tables 4, 5).


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CommentHabitaT: forest in hilly areas from sea level to at least 700 m in elevation. This species does quite well in disturbed forests and is commonly found on man-made structures. The holotype and paratypes from Kirirom National Park (FMNH 261851–54) were all found during the day (1400–1630 h) on or near the exterior walls of a building on a grassy plateau within open pine forest. FMNH 261851 was found on the branch of a large tree abutting an exterior wall, FMNH 261852 was found beneath a layer. of paint peeling off the exterior wall of the building approximately 2.5 m above the ground, and FMNH 261853–54 were found on the exterior walls. 
EtymologyThe specific epithet tokehos is the Khmer (Cambodian) word used for Ptychozoon. 
References
  • GRISMER, L. LEE; PERRY L. WOOD JR., JESSE L. GRISMER, EVAN S. H. QUAH, NEANG THY, SOMPHOUTHONE PHIMMACHAK, NIANE SIVONGXAY, SENGVILAY SEATEUN, BRYAN L. STUART, CAMERON B. SILER, DANIEL G. MULCAHY, TASHITSO ANAMZA & RAFE M. BROWN 2019. Geographic structure of genetic variation in the Parachute Gecko Ptychozoon lionotum Annandale, 1905 across Indochina and Sundaland with descriptions of three new species. Zootaxa 4638 (2): 151–198 - get paper here
  • Lalremsanga, Hmar Tlawmte; Lal Muansanga, Mathipi Vabeiryureilai & Zeeshan A. Mirza 2023. A new species of Parachute Gecko of the subgenus Ptychozoon (Sauria: Gekkonidae: Gekko) from the Indo-Burma region. Salamandra 59 (2): 125–135 - get paper here
  • Wood Jr, Perry L.; Xianguang Guo, Scott L. Travers, Yong-Chao Su, Karen V. Olson, Aaron M. Bauer, L. Lee Grismer, Cameron D. Siler, Robert G. Moyle, Michael J. Andersen, Rafe M. Brown 2019. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. bioRxiv 717520 [preprint] - get paper here
  • Wood, P. L., Guo, X., Travers, S. L., Su, Y. C., Olson, K. V., Bauer, A. M., Grismer, L. L., Siler, C. D., Moyle, R. G., Andersen, M. J. and Brown, R. M. 2020. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Molecular Phylogenetics and Evolution 146: 106731 [corrigendum: MPE 164: 107255] - get paper here
 
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