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Gloydius ussuriensis (EMELIANOV, 1929)

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Ussuri Mamushi
Russian: Уccурийский щитомордник
Chinese: 乌苏里蝮 
SynonymAncistrodon blomhoffi ussuriensis EMELIANOV 1929: 123
Agkistrodon halys ussuriensis — KLEMMER 1963: 400
Agkistrodon caliginosus GLOYD 1972: 563
Agkistrodon halys ussuriensis — HARDING & WELCH 1980
Gloydius caliginosus — HOGE & ROMANO-HOGE 1981: 195
Agkistrodon ussuriensis — TORIBA 1986
Gloydius ussuriensis — KRAUS MINK & BROWN 1996
Agkistrodon caliginosus — WELCH 1994: 11
Agkistrodon blomhoffi intermedius — WELCH 1994: 11
Agkistrodon ussuriensis — ORLOV & BARABANOV 1999
Gloydius ussuriensis — MCDIARMID, CAMPBELL & TOURÉ 1999: 309
Gloydius ussurriensis [sic] — ZHOU et al. 2001
Agkistrodon blomhoffi ussuriensis — TRUTNAU 2002
Gloydius ussuriensis — GUMPRECHT et al. 2004
Gloydius ussuriensis — WALLACH et al. 2014: 312
Gloydius ussuriensis — ORLOV et al. 2014 
DistributionE Russia

caliginosus (invalid): Korea, Manchuria; Type locality: vicinity of Seoul, Korea.

Type locality: Suchan River, Primorskiy Territory, Siberia (restricted by CHERNOV 1934); Terra typica restricta (hoc loco). Russian Far East, Primorskii Region, Tetyukhe river valley, Vladimiro-Monomakhovo village [ORLOV & BARABANOV 1999].  
Reproductionovovivparous 
TypesLectotype. ZISP 13327. Russian Far East, Primorskii Region, Tetyukhe river valley, Vladimi- ro-Monomakhovo village. Leg.: O. Djukin, June 20, 1909. 
DiagnosisDiagnosis: Total length up to 650 mm. Dorsal scales in 21 rows at midbody. Ventrals 139 – 161, subcaudals 34 – 54, supralabials 7.


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CommentVenomous!

Synonymy partly after KHALIKOV & ANANJEVA (pers. comm.)

Distribution: see map in ORLOV & BARABANOV 1999 and revised map in Orlov et al. 2014: Fig. 2 (which does not have ussuriensis for South Korea, North Korea,
NE China (from N Liaoning to Heilongjiang), so these are removed from its distribution. For a map of localities see Shi et al. 2021: 102 (Fig. 6). 
References
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