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Gloydius liupanensis LIU, SONG & LUO, 1989

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesChinese: 六盘山蝮 
SynonymGloydius halys liupanensis LIU, SONG & LUO 1989
Gloydius strauchi liupanensis — WANG & ZHAO 2006
Gloydius liupanensis — XU et al. 2012
Gloydius liupanensis — WANG et al. 2019 
DistributionChina (Gansu), 2300 m elevation

Type locality: Liupan Mountains in the Ningxia Huizo Region, east of the Qingyang area in der Province Gansu. Only known from the type locality.  
ReproductionViviparous. 
TypesType: unlocated fide McDIARMID et al. 1999: 308, given as Department of Biology, Lanzhou University, Nr. 85001, adult male; allotype given as 85004, adult female. 
DiagnosisDiagnosis: The new subspecies differs from the other 3 subspecies of our country in 1) difference between the females and the males in morpha; 2) body sides with the white lines from head to tail; 3) 19-21 rows of dorsal scales in the middle body; 4) ventrals + subcaudals 192—201 (on an average 196). (Liu et al. 1989, english abstract)


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CommentVenomous!

Synonymy: Zhao et al. (1998) synonymized Gloydius halys liupanensis with G. strauchi which was maintained by WALLACH et al. 2014: 311. GUMPRECHT et al. (2004) do not recognize the subspecies liupanensis, mogoi, and affinis. Zhao 2006 listed Gloydius halys cognatus, liupanensis, qinlingensis, and stejnegeri as synonyms of either G. strauchi or G. intermedius.

Phylogenetics: see XU et al. 2009.

Subspecies: Agkistrodon halys ussuriensis and other subspecies have been elevated to full species status.

Conservation: one of the 30 most endangered viper species (Maritz et al. 2016).

Distribution: for a map of localities see Shi et al. 2021: 102 (Fig. 6) 
EtymologyNamed after the type locality. 
References
  • Liu, N., Z. SONG & W. LUO 1989. A new subspecies of Agkistrodon halys. J. Lanzhou Univ. Nat. Sci. 25: 114-117
  • Maritz, Bryan; Johannes Penner, Marcio Martins, Jelka Crnobrnja-Isailović, Stephen Spear, Laura R.V. Alencar, Jesús Sigala-Rodriguez, Kevin Messenger, Rulon W. Clark, Pritpal Soorae, Luca Luiselli, Chris Jenkins, Harry W. Greene 2016. Identifying global priorities for the conservation of vipers. Biological Conservation, doi:10.1016/j.biocon.2016.05.004 - get paper here
  • Shi J-S, Liu J-C, Giri R, Owens JB, Santra V, Kuttalam S, Selvan M, Guo K-J, Malhotra A 2021. Molecular phylogenetic analysis of the genus Gloydius (Squamata, Viperidae, Crotalinae), with description of two new alpine species from Qinghai-Tibet Plateau, China. ZooKeys 1061: 87-108 - get paper here
  • Shi, Jingsong; Gang Wang; Xi’er Chen; Yihao Fang; Li Ding; Song Huang; Mian Hou, Jun Liu, and Pipeng Li 2017. A new moth-preying alpine pit viper species from Qinghai-Tibetan Plateau (Viperidae, Crotalinae). Amphibia-Reptilia, 38 (4): 517 – 532;<br>DOI: 10.1163/15685381-00003134 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Wang, Kai; Jinlong Ren, Wenjie Dong, Ke Jiang, Jingsong Shi, Cameron D. Siler and Jing Che 2019. A New Species of Plateau Pit Viper (Reptilia: Serpentes: Gloydius) from the Upper Lancang (= Mekong) Valley in the Hengduan Mountain Region, Tibet, China. Journal of Herpetology 53 (1): 224-236 - get paper here
  • Wang, Xiang-jun; Zhao, Er-mi. 2006. A preliminary taxonomic status of Gloydius qinlingensis and G. liupanensis. Sichuan Journal of Zoology 25 (2):210-213. [In Chinese]
  • XU, Yan; Qin LIU, Edward A MYERS, , Lian WANG, Song HUANG, Yun HE,Peihao PENG and Peng GUO 2012. Molecular Phylogeny of the Genus Gloydius (Serpentes: Crotalinae). Asian Herpetological Research 3 (2): 127–132 - get paper here
 
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