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Gloydius lateralis ZHANG, SHI, LI, YAN, WANG, DING, DU, PLENKOVIĆ-MORAJ, JIANG & SHI, 2022

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Jiuzhai pitviper
Chinese: 九寨蝮 (Jiǔ Zhài Fù) 
SynonymGloydius lateralis ZHANG, SHI, LI, YAN, WANG, DING, DU, PLENKOVIĆ-MORAJ, JIANG & SHI 2022
Gloydius strauchi – LI et al. 2004 
DistributionChina (Sichuan province)

Type locality: China, Sichuan province, Aba Tibetan and Qiang Autonomous Prefecture, Jiuzhaigou National Nature Reserve (33.26°N, 103.93°E), 2072 m elevation.  
Reproduction 
TypesHolotype: CIB 119377, аdult female, collected by C. L. Zhao, P. Yan, & T. Yang, on 8 June 2021.
Paratypes (n = 8): Three adult females (IVPP OV 2727, CIB 119378, and CIB 119379) collected by P. Yan and M. H. Zhang on 31 August 2021; four adult females (SAFS2021001–SAFS2021004) collected by P. Wang, on June 2021; one adult male (CIB 87280) collected by C. Li, on 23 May 2002. 
DiagnosisDiagnosis (and/or description): The above-mentioned specimens were identified as members of the genus Gloydius based on the small body size, bilateral pits, and divided subcaudal scales. G. lateralis sp. nov. differs from other congeneric species by a combination of the following characteristics: (1) relatively larger eyes (ED/HL: 0.145–0.171, n = 5); (2) three palatine teeth; (3) 20 or 21 rows of midbody dorsal scales; (4) ventrals 151–163 (n = 9); (5) subcaudals 38–49 (n = 6); (6) dorsal body laurel green or light brown with four rows of zigzag, dark brown patches, the medial two rows separated from each other by alternate phyllotaxis pattern in background color; (7) continuous, regular greyish-brown ventrolateral stripe on each side of body and tail. (Zhang et al. 2022)


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Comment 
EtymologyThe specific epithet lateralis refers to the unique continuous, regular, greyish-brown ventrolateral stripes at the junction of ventrals and the first row of dorsal scales. The common name refers to its type locality, Jiuzhaigou National Nature Reserve. 
References
  • Kwet, A. 2023. Liste der im Jahr 2022 neu beschriebenen Reptilien. Elaphe 2023 (3): 48-73
  • Li, C., Sun, Z.Y., Cai, Y.S., Liu, S.Y., Ran, J.H., Liu, Z.J., Wang, Y.Z. 2004. The Herpetofaunal Diversity in Jiuzhaigou National Nature Reserve, China. [in Chinese] Chinese Journal of Zoology, 39(2): 74–77
  • Zhang M-H, Shi S-C, Li C, Yan P, Wang P, Ding L, Du J, Plenković-Moraj A, Jiang J-P, Shi J-S 2022. Exploring cryptic biodiversity in a world heritage site: a new pitviper (Squamata, Viperidae, Crotalinae) from Jiuzhaigou, Aba, Sichuan, China. ZooKeys 1114: 59-76 - get paper here
 
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