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Gloydius lipipengi SHI, LIU & MALHOTRA, 2021

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Nujiang pit viper
Chinese: 怒江蝮 (Nù Jiāng Fù) 
SynonymGloydius lipipengi SHI, LIU & MALHOTRA in SHI et al. 2021 
DistributionZayu, Tibet, China

Type locality: Zawalong, Zayu, Tibet, China  
Reproduction 
TypesHolotype. IVPP OV2720 (G2, Figs 1–4), adult male, collected from Muza Village, Zayu, Nyingchi Prefecture, Tibet (2883 m), by Jin-Cheng Liu, on 8 September 2014. 
DiagnosisDiagnosis: The specimens of the new species, IVPP OV 2720, IVPP OV 2725 and IVPP OV 2726 were identified as the member of the genus Gloydius based on the small body size, bilateral pits, and divided subcaudal scales (Hoge and Romano- Hoge 1981). Gloydius lipipengi sp. nov. differs from other congeneric species in the following characteristics: i) third supralabial scale not touching the orbit; ii) a pair of prominent black markings on the occiput; iii) black-bordered greyish cheek stripe extending from the posterior margin of orbit (not separated by the postoculars) to the ventral surface of the neck; iv) black irregular annular crossbands on the mid-body; iv) two rows of black blotches on the ventral side; v) 23-21-15 circum-body scales; vi) 165 ventral scales; and vii) 46 subcaudal scales. (Shi et al. 2021).


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CommentDistribution: for a map of localities see Shi et al. 2021: 102 (Fig. 6) 
EtymologyThe specific epithet of the new species from Tibet is dedicated to the senior author’s Master’s supervisor, Professor Pi-Peng Li (Institute of Herpetology, Shenyang Normal University) on Li’s sixtieth birthday. Prof. Li has devoted himself to the study of the herpetological diversity of the Qinghai-Tibet Plateau. The senior author became an Asian pit viper enthusiast and professional herpetological researcher under his instruction. 
References
  • Shi J-S, Liu J-C, Giri R, Owens JB, Santra V, Kuttalam S, Selvan M, Guo K-J, Malhotra A 2021. Molecular phylogenetic analysis of the genus Gloydius (Squamata, Viperidae, Crotalinae), with description of two new alpine species from Qinghai-Tibet Plateau, China. ZooKeys 1061: 87-108 - get paper here
 
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