Gloydius brevicaudus (STEJNEGER, 1907)

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Higher Taxa	Viperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies	Gloydius brevicaudus brevicaudus (STEJNEGER 1907) Gloydius brevicaudus siniticus (GLOYD 1977)
Common Names	G: Kurzschwanz-Mamushi Chinese: 短尾蝮
Synonym	Agkistrodon blomhoffii brevicaudus STEJNEGER 1907: 463 Ancistrodon halys brevicaudus NIKOLSKY 1916 Ancistrodon halys brevicaudus — RENDAHL 1933: 21 Ancistrodon halys brevicaudus — SHANNON 1956 Gloydius blomhoffii brevicaudus — HOGE & ROMANO-HOGE 1981: 195 Agkistrodon blomhoffii brevicaudus — WELCH 1994: 11 Agkistrodon blomhoffii dubitatus GLOYD 1977 Agkistrodon blomhoffii brevicaudus — TRUTNAU 2002 Gloydius brevicaudus — ZHOU et al. 2001 Gloydius brevicaudus — WANG & ZHAO 2007 Gloydius brevicaudus — GAULKE 2008 Gloydius brevicaudus — WALLACH et al. 2014: 310 Gloydius brevicaudus — ORLOV et al. 2014 Gloydius brevicaudus brevicaudus — DAVID & VOGEL 2015 Gloydius brevicaudus siniticus (GLOYD 1977) Agkistrodon blomhoffii siniticus GLOYD 1977 Gloydius blomhoffii siniticus — HOGE & ROMANO-HOGE 1981 [1979] Gloydius blomhoffii siniticus — ORLOV & BARABANOV 1999 Gloydius blomhoffii siniticus — GUMPRECHT et al. 2004 Gloydius brevicaudus siniticus — WANG & ZHAO 2007 Gloydius blomhoffii siniticus — ORLOV et al. 2014 Gloydius brevicaudus siniticus — DAVID & VOGEL 2015
Distribution	North Korea, South Korea, China (Liaoning, Jilin, Guangdong: HR 33: 226); Russia (Pozharsky District, Primorsky Krai) [iNaturalist] Type locality: Fusan = (Pusan), S. Korea. siniticus: China (Yangtze River basin from E Sichuan to the delta; north to Anhwei, Kiangsu and Shantung; south into N Hunan and Kiangsi). Type locality: Ningkwo (= Ningguo Co.), Anhwei (= Anhui) Province, China.
Reproduction	viviparous
Types	Holotype: USNM 17507 Holotype: AMNH 25554 [siniticus]
Diagnosis
Comment	Venomous! Synonymy: G. blomhoffi dubitatus was listed as a synonym of G. blomhoffi by McDiarmid et al. 1999 but as a synonym of G. brevicaudus by Zhao and Adler (1993), David and Ineich (1999), and Orlov et al. 2014. Some of these authors also consider A. b. siniticus to be a junior synonym of Gloydius brevicaudus although it has been listed as valid more recenlty. Distribution: see map in ORLOV & BARABANOV 1999. G. b. brevicaudus and G. b. siniticus are allopatric with a large gap in between their reanges.
References	Chen, Y.; Zhang, D.; Jiang, K. & Wang, Z. 1992. Evaluation of Snake Venoms among Agkistrodon Species in China. Asiatic Herpetological Research 4: 58-61 - get paper here David, P. & Vogel, G. 2015. An updated list of Asian pitvipers and a selection of recent publications, in: Visser: Asian Pitvipers. Breeding Experience & Wildlife. Edition Chimaira, Frankfurt am Main, 571pp. ISBN 978-3-89973-450-8 - get paper here Gaulke, M. 2008. Der Avilon Montalban Zoological Park, Philippinen. Reptilia (Münster) 13 (69): 76-81 - get paper here Gloyd, H.K. 1977. Descriptions of new taxa of crotalid snakes from China and Ceylon (Sri Lanka). Proc. Biol. Soc. Washington 90: 1002-1015 - get paper here Guo, P. 2000. The anatomy of the skeletal system of Gloydius brevicaudus. Cultum Herpetological Sinica, Zunyi 8:34~38 Guo, P.; Zhang, F.J. & Chen, Y.Y. 1999. The Hemipenes of Chinese Species of Deinagkistrodon and Gloydius (Serpentes: Crotaiinae). Asiatic Herpetological Research 8: 38-42 - get paper here Hoge & Romano-Hoge 1981. Poisonous snakes of the world. Part I. Check list of the pit vipers, Viperoidea, Viperidae, Crotalinae. Mem. Inst. Butantan 42/43 [1978-1979]: 179-309 - get paper here Huang, M.; Cao, Y.; Zhu, F. & Qu, Y. 1992. Female Reproductive Cycle and Embryonic Development of the Chinese Mamushi (Agkistrodon blomhoffii brevicaudus). Asiatic Herpetological Research 4: 62-67 - get paper here Koo, Kyo Soung 2019. First Report on the Hybridization between Gloydius brevicaudus and G. intermedius. Herpetology Notes 12: - get paper here Nikolsky, A. M. 1916. Faune de la Russie, Reptiles Vol. 2, Ophidia. Petrograd, 396 pp. [English translation by A. Mercado, Israel and Smithsonian Institution, 247 pp.] Orlov, Nikolai L. and Andrei V. Barabanov. 1999. Analysis of nomenclature, classification, and distribution of the Agkistrodon halys - Agkistrodon intermedius complexes: a critical review. Russ. J. Herpetol. 6 (3): 167-192 - get paper here Orlov, Nikolai L.; Yuri Sundukov, Ivan Kropachev 2014. Distribution of Pitvipers of «Gloydius blomhoffii» Complex in Russia with the First Records of Gloydius blomhoffii blomhoffii at Kunashir Island (Kuril Archipelago, Russian Far East). Russ. J. Herpetol. 21 (3): 169-178 - get paper here Rendahl, Hialmar 1933. Die Unterarten des Ancistrodon halys Pall. nebst elnigen Bemerkungen zur Herpetologie Zentralasiens. Arkiv för Zoologi 25 (8): 1-31 Seung Hoon, Cha 2012. Snake, the world most beautifull curve [in Korean]. Hownext, 304 pp. [ISBN 978-89-965656-7-3] - get paper here Shannon, F.A. 1956. The reptiles and amphibians of Korea. Herpetologica 12 (1): 22-49 - get paper here Stejneger, LEONHARD H. 1907. Herpetology of Japan and adjacent territory. Bull. US Natl. Mus. 58: xx, 1-577 - get paper here Trutnau, L. 2002. Die Kurzschwanz-Mamushi Agkistrodon blomhoffii brevicaudus STEJNEGER 1907 - eine selten gepflegte ostasiatische Grubenotter. Herpetofauna 24 (138): 19-28 - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here Welch, K. R. G. 1994. Snakes of the World. A Checklist. I. Venomous snakes. KCM Books, Somerset, England. XIONG Ye, DING Li 2012. A Taxonomic Status of Fu Snakes in Mt. Wuling, Xinglong County, Hebei Province. Sichuan Journal of Zoology 31 (5) - get paper here XU, Yan; Qin LIU, Edward A MYERS, , Lian WANG, Song HUANG, Yun HE,Peihao PENG and Peng GUO 2012. Molecular Phylogeny of the Genus Gloydius (Serpentes: Crotalinae). Asian Herpetological Research 3 (2): 127–132 - get paper here Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp. Zhou, Ji-Liang; Zhang, Ya-Ping; Huang, Mei-Hua; Chen, Yong-Jiu; Chen, Xiao-Qing and Yao, Geng-Dong 2001. Phylogenetic relationships among crotalinae based on mitochondrial cytochrome B gene sequence variations [in Chinese]. Acta Zool. Sinica 47 (4): 361-366
External links	National Center for Biotechnology Information http://www.airiti.com/ceps/ec_en/ecjnlarticleView.aspx?jnlcattype=1&jnlptype=5&jnltype=42&jnliid=3001&issueiid=51603&atliid=849419 Google images

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