Gloydius intermedius (STRAUCH, 1868)
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Higher Taxa | Viperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) |
Subspecies | |
Common Names | E: Central Asian pitviper Russian: Средний щитомордник Chinese: 中介蝮 |
Synonym | Trigonocephalus intermedius STRAUCH 1868: 295 Agkistrodon halys intermedius STRAUCH 1876 Halys intermedia — PETERS 1877: 735 Ancistrodon intermedius — BOULENGER 1890: 140 Ancistrodon intermedius — BOULENGER 1896: 525 Agkistrodon blomhoffi intermedius — STEJNEGER 1907: 464 Ancistrodon blomhoffi intermedius — DESPAX 1913: 181 Ancistrodon halys intermedius — EMELIANOV 1929: 130 Ancistrodon halys intermedius — RENDAHL 1933: 15 Ancistrodon saxatilis EMELIANOV 1937 Agkistrodon saxatilis GLOYD 1972 Agkistrodon halys cognatus GLOYD 1977 Agkistrodon halys intermedius — HARDING & WELCH 1980 Agkistrodon shedaoensis continentalis ZHAO 1980: 12 Agkistrodon saxatilis — SCHNEEWEISS 1989 Agkistrodon intermedius saxatilis — WELCH 1994: 13 Agkistrodon intermedius intermedius — WELCH 1994: 11 Gloydius saxatilis changdaoensis LI 1999 Agkistrodon intermedius — ORLOV & BARABANOV 1999 Gloydius saxatilis — MCDIARMID, CAMPBELL & TOURÉ 1999: 307 Gloydius intermedius — MCDIARMID, CAMPBELL & TOURÉ 1999: 306 Gloydius intermedius — GUMPRECHT et al. 2004 Gloydius saxatilis — JIANG & ZHAO 2009 Gloydius intermedius — WALLACH et al. 2014: 311 Gloydius saxatilis — WALLACH et al. 2014: 311 Gloydius intermedius — DAVID & VOGEL 2015 Gloydius intermedius — WAGNER et al. 2015 |
Distribution | N China (incl. Liaoning, Jilin, Heilongjiang), E Russia, North Korea, South Korea Type locality: Irkutsk, Eastern Siberia; Terra typica restricta (hoc loco). Malyi Hinggan Mountain Range (Xiao Hinggan Ling), Cape Tyr on the Amur River and Cape Kofel on the Ussuri River, Russian Far East Region. saxatilis: NE China south to Shandong, Russia (east of Selemdzha River in E Siberia, Primorskye Kray and Khabarovsk Kray, Far East), Korea; Type locality: Vladivostok River, Russia changdaoensis: China (Daheishan Is., Changdao County; Zifu Peninsula and Jia Is., Yantai City, Shandong Province); Type locality: Daheishan Island, Changdao County, Shandong Province, China. |
Reproduction | viviparous |
Types | Lectotype. ZISP 2221. Prom. Tyr (Amur). Leg.: Schrenck, 1855 Syntypes: ZIAS 9456, 13291–92 (Vladivostok) and ZIAS 13308 (Suchan); of the many specimens listed in the original description (Emelianov, 1937:27 and Tables 1-3), only two are consistent with the type locality as given in his English summary: Table II, No. 17, D(al'nii). V(ostok). (= Far East U(niversitet)., r. Suchan, 8 VIII 1927, (coIl.) Gasovskii, and Table III, No.7, Zoological Cabinet. GDU, vic. Vladivostok, 30 IX 1927, (coIl.) A. A. Emelianov. Gloyd and Conant (1990:358) seemingly were unsure of which specimens were part of type series. Holotype: DLNM (?), Natural History Museum of Snake Island, Dalian [changdaoensis] |
Diagnosis | Diagnosis. Total length up to 800 mm and more. Dorsal scales in 23 (rarely 21) rows at midbody. Supralabials 7 – 8; ventrals 148 – 175; subcaudals 34–52. Diagnosis. A large viper of the genus, up to at least 800 mm total length. Supralabial scales 7-8. Dorsal scales in 23, very rarely 21, rows around midbody; ventral scales between 148- 175; subcaudal scales paired, between 34-52. Body with 28-45 dark transverse bands, each 3- 6 scales wide and extending down to scale row 2 or 1 [WAGNER et al. 2015]. |
Comment | Venomous! Synonymy partly after KHALIKOV & ANANJEVA (pers. comm.). G. saxatilis is not recognized by GUMPRECHT et al. 2004. ORLOV & BARABANOV 1999 synonymized G. saxatilis with G. intermedius. See Jiang & Zhao 2009 for comparisons of G. saxatilis, shedaoensis, and their new species lijianlii. Distribution: see map in ORLOV & BARABANOV 1999. NCBI taxonID: 92067 [saxatilis] |
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