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Goggia braacki (GOOD, BAUER & BRANCH, 1996)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Braack’s Dwarf Leaf-Toed Gecko, Karoo Leaf-Toed Gecko 
SynonymPhyllodactylus braacki GOOD, BAUER & BRANCH 1996
Goggia braacki — BAUER et al. 1997
Goggia braacki — RÖSLER 2000: 83
Goggia braacki — BATES et al. 2014:
Goggia braacki — BATES et al. 2014: 108
Goggia braacki — TELFORD et al 2022 
DistributionRepublic of South Africa (Karoo National Park)

Type locality: Nuweveld escarpment near radio mast, Karoo National Park, Western Cape Province [then Cape Province],
South Africa (32°16' S, 22°30'E; 3222BC), approx. 1800
m elevation.  
Reproductionoviparous 
TypesHolotype: PEM R4361, adult male Paratypes (10 specimens): CAS 199989-91, LSUM 57330-2, PEM R1!890, Rll922, R12379-80, Mountain View, 4.9 km from Puttersvlei Gate, Karoo National Park, Western Cape Province, South Africa (3222BA), 1790 m, all collected 10 September 1995 by W.R. Branch, A.M. Bauer, and D.A. Good. 
DiagnosisDiagnosis: A large species of the Phyllodactylus linealus complex, adult snout-vent length (SVL) 27-35 mm. Distinguished from other species in the genus and in the species complex by the following combination of characters: head depressed; dorsal body scalation homogeneous; midbody scale rows 79-88; 4 preanal pores in males; usually one nasorostral scale; granules across crown between orbits usually 15-17; dorsal coloration pattern dominated by a scalloped or reticulate pattern. Phyllodactylus braacki differs from P. microlepidolus, P. porphyreus, and P. swartbergensis by its smaller size, and from the last species and P. peringueyi by its homogeneous body scalation. It may be distinguished from P. linealus and P. gemmulus by its higher number of scales around mid-body (usually more than 80 vs. usually less than 80). It is distinguished from P. linealus and P. hexaporus by the presence of four (versus 5-7) preanal pores in males. The presence of a single nasorostral granule (usually) and a greater body size (maximum SVL > 32 mm) serve to distinguish P. braacki from the vel)' similar species P. essexi and P. rupicolus. Phyllodactylus braacki is morphologically indistinguishable from P. hewitti, from which it is geographically isolated, but may be diagnosed from this species on the basis of three uniquely occurring allozyme alleles (Good et al. 1996: 51).


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Comment 
EtymologyThe specific epithet is a patronym honoring Mr. Harold Braack, formerly warden of the Karoo National Park. 
References
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer A M. Good D A. Branch W R. 1997. The taxonomy of the Southern African leaf-toed geckos (Squamata: Gekkonidae), with a review of Old World "Phyllodactylus" and the description of five new genera. Proc. Cal. Acad. Sci. 49 (14): 447-497. - get paper here
  • CONRADIE, WERNER; WILLIAM R. BRANCH, & GILLIAN WATSON 2019. Type specimens in the Port Elizabeth Museum, South Africa, including the historically important Albany Museum collection. Part 2: Reptiles (Squamata). Zootaxa 4576 (1): 001–045 - get paper here
  • Good, D. A., A. M. BAUER & W. R. BRANCH 1996. A new species of Phyllodactylus (Squamata: Gekkonidae) from the Karoo National Park, South Africa. African Journal of Herpetology 45 (2): 49-58 [1997] - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Telford, N. S., ALEXANDER, G. J., BECKER, F. S., CONRADIE, W., JORDAAN, A., KEMP, L., ... & TOLLEY, K. A. 2022. EXTENSIONS TO THE KNOWN GEOGRAPHIC DISTRIBUTIONS OF REPTILES IN THE GREAT KAROO, SOUTH AFRICA. Herpetological Conservation and Biology, 17(1), 145-154 - get paper here
 
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