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Goggia lineata (GRAY, 1838)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Striped Dwarf Leaf-toed Gecko 
SynonymPhyllodactylus lineatus GRAY 1838: 268
Diplodactylus lineatus — GRAY 1845: 150
Phyllodactylus lineatus — GRAY 1849
Phyllodactylus lineatus — BOULENGER 1885: 92
Phyllodactylus porphyreus — WERNER 1910: 307 (part.)
Phyllodactylus lineatus lineatus — HEWITT 1937
Phyllodactylus lineatus — WERMUTH 1965: 138
Phyllodactylus lineatus — KLUGE 1993
Phyllodactylus lineatus — RÖSLER 1995: 146
Goggia lineata — BAUER et al. 1997
Goggia lineata — RÖSLER 2000: 83
Goggia lineata — BATES et al. 2014: 111 
DistributionRepublic of South Africa (Cape Province, S Warmbad),
S Namibia

Type locality: Namaqualand; Cape of Good Hope (fide BAUER et al. 1995).  
TypesSyntype: BMNH xxi.11.2-3 (fide Loveridge 1947: 250, De Lisle et al. 2013: 101), maybe lost (Heinicke et al. 2017) 
DiagnosisDiagnosis: Distinguished from other members of the P. lineatus complex by a combination of lhe following features: usually five preanal pores in males, a low number of midbody scale rows (64-84), few scales between the nostril and anterior margin of the orbit (7-10), a deep head, and a cylindrical body. In addition, it differs by 4-8 fixed alleles from the other taxa examined electrophoretically (Branch et al. 1995).

Description: (Figure IF, IG) Head deep, not dorsoventrally flattened (depth 1.53 times width; range 1.22 - 2.19); snout rounded, and about 1.6 - 2.0 times the eye diameter; ear opening small, obliquely rounded, and without eularged lobules or a tympanic shield; rostral subpentagonal with a median cleft above; nostril pierced between rostral, first supralabial and three nasals, the largest bordering the rostral; usnally I (49.8%) or 2 (43.5%) nasorostral granules, rarely 0 (1.4%) or 3 (5.5%); supralabials usually 7 (50.6%), often 6 (35.0%), rarely 8 (13.6%) or 9 (0.8%); infralabials usnally 6 (63.6%), often 7 (23.6%), rarely 5 (10.0%) or 8 (2.8%); mental subpentagular bordered by 2-3 (average 2.21), rarely 4, chinshields that are larger than adjacent granules; 5-9 (average 6.1) granules bordering chinshields. Body short (maximum snout-vent length 31.75 mm) and cylindrical, covered dorsally with uniform, smooth, flattened, subimbricate granules; belly covered in smooth, hexagonal, imbricate sc ales that are much larger than on back and may have denticulate edges. Limbs short to moderate and covered in uniform, flattened, subimbricate granules; digits flattened basally, with three rows of subdigital granules, the median series slightly broader; toe tips rounded, only slightly expanded and bearing a pair of large, subrectangular scansors that enclose a small claw. An angular series of preanal pores present only in males, usnally 5 (87.1%), rarely 4 (6.45%) or 6 (6.45%); two to three enlarged tubercular scales on either side of tail base near vent, that are larger in males. T ail cylindrical, tapering, and, when umegenerated, usually longer (up to 1.3 times) than snout-vent length; covered above with regular rows of uniform, smooth granules, that are much larger, flattened and imbricate on ventral surface (Branch et al. 1995).

Colour: Above grey, sometimes with brownish infusion, and with dark gray to black markings that may occur in two phases (intermediates may also occur). Striped phase (PEM R7607, Richtersveld National Park) -- a black line extends from the snout, through the middle of the eye and temporal region, on to the flanks and tail base; a pair of dorso-lateral lines originate on the sides of the crown, and fuse in the sacral region to extend on to the proximal region of the tail. Scalloped phase (PEM RII161, Schaapen Island, Langebaan) -- the dorsum is covered with an irregular series of pale dorsal spots, each emargined anteriorly by a black edge; these margins may fuse (particularly on the neck, back of the head, and sacral region) to form a scalloped pattern that on the proximal region of an original tail forms irregular bars; a faint stripe may occur on the lower flank, extending through the eye to the snout. The throat, ventrum and lower surfaces of the tail and limbs are cream in both phases. The distal region of the original tail, particularly in hatchlings, may be pink-brown (Branch et al. 1995). 
CommentSubspecies: Phyllodactylus lineatus essexi and P. l. rupicolus have been raised to species status by BAUER et al. 1997.

Type Species: Phyllodactylus lineatus GRAY 1838 is the type species of the genus Goggia BAUER et al. 1997. 
EtymologyNamed after Latin “linea”, meaning stripe or line. The genus Goggia has been named after the Afrikaans word “gogga”, meaning a “creepy-crawly”, usually an arthropod or reptile. 
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer A M. Good D A. Branch W R. 1997. The taxonomy of the Southern African leaf-toed geckos (Squamata: Gekkonidae), with a review of Old World "Phyllodactylus" and the description of five new genera. Proc. Cal. Acad. Sci. 49 (14): 447-497. - get paper here
  • Bauer, A.M., and Branch, W.R. 2003. The herpetofauna of the Richtersveld National Park, Northern Cape Province, Republic of South Africa. Herpetological Natural History 8: 111-160 [2001]
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Branch, W. R., A. M. BAUER & D. A. GOOD 1995. Species limits in the Phyllodactylus lineatus complex (Reptilia: Gekkonidae), with the elevation of two taxa to specific status and the description of two new species. J. Herp. Assoc. Africa 44 (2): 33-54. - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Branch, William R. 1994. Herpetofauna of the Sperrgebeit [sic] Region of Southern Namibia. Herpetological Natural History 2 (1): 1-11.
  • Gray, J. E. 1838. (A new species of lizard and land shells from South Africa.] Pp. 268-269. In Alexander, J. E., An expedition of discovery into the interior of Africa, through the hitherto undescribed countries of the Great Namaquas, Boschmans, and Hill Damaras, perf H. Colburn, London - get paper here
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • HEINICKE, MATTHEW P.; DILARA TURK, AARON M. BAUER 2017. Molecular phylogeny reveals strong biogeographic signal and two new species in a Cape Biodiversity Hotspot endemic mini-radiation, the pygmy geckos (Gekkonidae: Goggia). Zootaxa 4312 (3): 449–470 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Hewitt, J. 1937. A guide to the vertebrate fauna of the Eastern Cape Province, South Africa, Part II: reptile, amphibians, and freshwater fishes. Grahamstown, vii + 141 pp.
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp.
  • Werner,F. 1910. Reptilia et Amphibia. In Schultze, L., Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Südafrika. Band IV, Systematik und Tiergeographie Vertebrata B. Denkschr. Med.-Nat. Wiss. GeselI. Jena 16: 279-370 [1910] - get paper here
  • WITBERG, M. 2012. The herpetofauna of Schaapen Island, Langebaan, South Africa. African Herp News (56): 11-16 - get paper here
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