You are here » home › advanced search › search results › Gonatodes humeralis

Gonatodes humeralis (GUICHENOT, 1855)

iNaturalist logo

Can you confirm these amateur observations of Gonatodes humeralis?

Add your own observation of
Gonatodes humeralis »

Find more photos by Google images search: Google images

Higher TaxaSphaerodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: South American Clawed Gecko, Trinidad Gecko
S: Limpiacasa
Portuguese: Lagartinho-Bicudo, Lagartinho-Colorido, Lagartinho-Pintado, Lagartixa, Lagartixa-da-Mata, Lagartixinha-Amazônica, Lagarto, Lagarto-do-Folhiço 
SynonymGymnodactylus timoriensis DUMÉRIL & BIBRON 1836: 411
Gymnodactylus humeralis GUICHENOT 1855: 13
Gonatodes ferrugineus COPE 1864
Gymnodactylus incertus PETERS 1871: 397
Goniodactylus sulcatus O’SHAUGHNESSY 1876: 265
Gonatodes humeralis — BOULENGER 1885: 62
Gonatodes ferrugineus — BOULENGER 1885: 56
Gonatodes timorensis — BOULENGER 1885: 63 (nom. subst.)
Gonatodes timorensis — DE ROOIJ 1915: 25
Gonatodes humeralis — BURT & MYERS 1942
Cnemaspis timoriensis — WERMUTH 1965: 17
Cnemaspis timoriensis — KLUGE 1993
Gonatodes humeralis — KLUGE 1993
Gonatodes humeralis — GORZULA & SEÑARIS 1999
Cnemaspis (Cnemaspis) timoriensis — RÖSLER 2000: 63
Gonatodes humeralis — LEHR 2002: 72
Gonatodes humeralis — LIVIGNI 2013: 341
Cnemaspis timoriensis — RÖSLER 2016: 20
Gonatodes ferrugineus — PINTO et al. 2018
Gonatodes humeralis — PINTO et al. 2018
Gonatodes ferrugineus — AUGUSTE 2019
Gonatodes ferrugineus — RIVAS et al. 2021 
DistributionTrinidad, French Guiana, Suriname, Ecuador,
Guyana, Peru (Loreto), Bolivia (Beni, Pando, Santa Cruz), NE Venezuela, Colombia, Brazil (Pará, Acre, Amazonas, Rondônia, Piauí etc.)

Type locality: “Pebas” [Hochland von Peru] [Gymnodactylus incertus PETERS 1871]

ferrugineus: Trinidad  
Reproductionoviparous 
TypesHolotype: lost; (was ZMB 7189, but lost fide BAUER et al. 1995); also given as USNM (presumably also lost fide Rivero-Blanco 1979, Pinto et al. 2018)
Holotype: MNHN-RA 810; the other known specimen assigned to this name is ZMB 9478 (designated as paratype by Bauer & Günther 1991 but not valid following Bour & Brygoo 2013 because the species was described based on a single specimen). [timoriensis] 
DiagnosisDiagnosis. Gonatodes with proximal subdigital lamellae as wide as digit, in total 15-21 under fourth toe. Two lateral rows of scales at each side on distal part of fingers and toes. Tail ventrally with a repetitive sequence of two single midventrals (one after the other), each in contact latero-distally with one scale per side, followed by a slightly larger, single midventral in contact at each side (latero-distally) with two scales. Scales around midbody 100-137. Ventrals 48-78. Male, in life, with a red, yellow and brown vermiculated dorsal pattern; head with pale grey or yellow, and red stripes and spots; a white or yellow antehumeral bar is present, preceded by a round black spot (Avila-Pires 1995: 273 ff).

Description. Sphaerodactyline gecko with maximum SVL in males of 41.5 mm (MPEG 15284), in females of 40.5 mm (MPEG 15218). Head 0.21-0.28 (n= 111) times SVL, proportionally smaller in larger specimens; 1.4-1.8 (1.60 ± 0.08, n= 110) times as long as wide; 1.1-1.7 (1.32 ± 0.11, n= 110) times as wide as high. Snout round, moderately elongate, gently sloping toward top of head. Neck slightly narrower than head and body. Body cylindrical. Limbs well developed, forelimbs 0.29-0.41 (0.34 ± 0.02, n= 86) times SVL, hind limbs 0.38-0.52 (0.44 ± 0.03, n= 72) times. Tail round in cross section, tapering toward tip, 1.1-1.4 (1.30 ± 0.07, n= 29) times SVL.
Tongue relatively wide, slightly narrowing anteriorly, with a round tip; covered anteriorly by scalelike papillae, posteriorly villose; tip with a short median cleft. Teeth small, conical, subequal.
Rostral large, convex, distinctly visible from above, with a long median cleft extending from posterior margin. Postrostrals 2-6, mostly three, lateral ones (supra- nasals) much larger than median ones, median ones about as large as adjacent scales on snout. Nostril bordered by rostral, first supralabial, three postnasals and lateral postrostral (supranasal). Postnasals as large as, or slightly larger than, adjacent loreals. Scales on snout and on loreal region round to polygonal, slightly convex, juxtaposed. Loreal scales 7-12 (mostly 8-10) in a line between postnasals and anterior margin of orbit. From between eyes toward posterior part of head the scales decrease in size and become granular. Scales on supraorbital region similar to, and continuous with those on top of head. Supraciliary flap with a smooth margin, or with up to seven slightly enlarged, prominent scales on anterior upper margin, of which the posterior ones may form small spines over the eye. Pupil round. Supralabials 5-9, decreasing in size posteriorly; 4-7 to below centre of eye. Scales on temporal region small, granular, like those on posterior upper part of head. Ear-opening much smaller than eye, round to vertically oval or triangular.
Mental large, roughly trapezoid with convex anterior border, to rhomboid. Two, rarely three, postmentals. Scales on anterior part of chin polygonal, gradually decreasing in size from postmentals; posteriorly round, relatively small; all juxtaposed. Infralabials 5-8, decreasing in size posteriorly; 3-6 to below centre of eye.
Scales on nape and sides of neck granular, slightly larger posteriorly. Scales on throat smooth, imbricate, with round posterior margin, with a short transitional zone with the granular scales on chin.
Dorsals granular, slightly larger than scales on top of head. Scales on flanks mostly like the dorsals, ventrolaterally grading into ventrals. Ventral region with scales distinctly larger than dorsals, smooth, hexagonal, imbricate, in oblique rows; 48-78 (61.4 ± 4.7, n= 109) scales along the midventral line between anterior level of forelimbs and vent. Scales around midbody 100-137 (117.4 ± 6.6, n= 107). Scales on preanal plate similar to ventrals, except for border of vent, which has very small scales. No preanal or femoral pores. Males with an escutcheon area covering posterior part of belly, and four to five rows of scales on ventral surface of thighs.
Scales on dorsal surface of tail smooth, roundish, imbricate, on the sides increasing in size toward ventral surface. A midventral row of transversely enlarged scales, with the sequence l ' l ' i " (figs. 2, 89; some variation exists, but this sequence usually predominates).
Scales on anterodorsal surface of forelimbs, anterior and ventral surfaces of thighs and ventral surface of lower legs smooth, roundish, imbricate; on forelimbs similar in size to scales on dorsal part of tail, on hind limbs increasing in size toward ventral surface; on other surfaces of both fore- and hind limbs, scales granular. Lamellae under third finger 12-17 (14.7 ± 0.8, n= 217), of which four, rarely three or five, basal ones distinctly larger and flattened; under fourth finger 12-18 (15.7 ± 0.9, n= 217), with five, occasionally four or six, enlarged basal ones; under fourth toe 15-21 (17.7 ± 1.0, n= 211), with six or seven, occasionally five or eight, enlarged basal ones. Fingers and toes with two lateral rows of scales distally. Claws exposed, non retractile, between two basal scales (Avila-Pires 1995: 273 ff).

Sexual dichromatism evident, with males much more colourful. In life, males dorsally with head variably ornamented on snout and top of head with pale or light neutral grey (85, 86), olive-grey (42), or yellow (spectrum-yellow, 55; sulphur-yellow, 157; yellow-green, 58); red (crimson, 108; poppy-red, 108A; scarlet, 14; geranium-pink, 13); and olive or brown (Prout's brown, 121A; hair-brown, 119A). A frequent pattern is that of a large light (grey, yellow) spot on snout, a rhomboid light spot on top of head, and a light arc posteriorly, at each side starting at posterior corner of eye, in some specimens continuing also anterior to eyes; between the light areas, predominantly red (especially on the snout), or red and brown. Nape and sides of head and neck yellow, red, and olive or brown. Antehumeral bar usually spectrum-yellow, occasionally greenish-yellow or whitish, preceded by a round black spot (a second, smaller, black spot may also be present dorsally). Back from predominantly brown or olive, to densely vermiculated with yellow, or yellow and red. Flanks similar to the back, or with the yellow vermiculation turning into white. Four longitudinal rows of round spots may be present along body, either all four with red spots, or the two lateral ones with black spots; in RMNH 26408 there were two series of paravertebral pale brown spots, more vivid at base of tail (tail regenerated posteriorly). Limbs coloured like body. Dorsal surface of tail, in MPEG 15871, olive (30) with smoke-grey (45) spots at base, distally natal-brown (219A) and sepia (219), separated by narrow beige (219D) rings; in MPEG 16013 proximal part of tail marbled brown, distal part, which is regenerating, marbled brown and pale flesh-colour. Ventrally, head spectrum-yellow (55) or sulphur-yellow (57); belly lime-green (59), smoke-grey (44, 45), drab-grey (119D), or anteriorly white peppered with brown, posteriorly straw-yellow (56); escutcheon area similar in colour to belly, or yellowish, yellow-ocher (123C); tail spectrum-orange (17), salmon (106), flesh-ocher (132D), or pale flesh colour (5), in some specimens marbled (along all of it or posteriorly) with light grey or natal-brown (219A). Iris brown with a narrow orange-brown rim around the pupil. Tongue white with grey tip.
In females, dorsal region olive-brown (28) with black spots and smoke-grey (44) stripes (RMNH 26410); with a cinnamon-drab (219C) vertebral band bordered by sepia (219), flanks vandyke-brown (121) with irregular sepia spots (MPEG 15880); a mixture of brown and olive-brown, with a drab (27) vertebral band (RMNH 26407); or back smoke-grey (45) with brownish-olive (29) markings, and flanks olive (30) with smoke-grey (44) markings (RMNH 26416). A cream antehumeral bar, preceded by an olive-brownspot may be present (MPEG 15913). Tail on dorsal surface cinnamon-brown (33) with cinnamon (39) and warm-sepia (221A) spots (RMNH 26410), predominantly sepia (219) and mikado-brown (121C) (MPEG 15880), or flesh-ocher (132D) and dark brown (MPEG 16014). Ventrally, head smoke-grey (44) or cream colour (54), with or without olive-brown or greyish-brown spots; belly smoke-grey (44), cream colour (54), sulphur-yellow (157), or chamois (123D); tail spectrum-orange (17), flesh-ocher (132D), light-russet-vinaceous (221D), salmon (106), or beige (219D) and pale brown. Iris and tongue like in males (Avila-Pires 1995: 273 ff).

Coloration in preservative, males with a characteristically antehumeral white bar, almost reaching the middorsal region and usually preceded by an oval, black spot; in some specimens, the black spot is preceded, in its turn, by another (shorter) white bar; in other specimens, a second, smaller black spot is present at the level of the dorsal end of the antehumeral bar. Head and body dorsally with a finer or coarser dark and light brown vermiculation; a "U"-shaped white band on posterior upper part of head, connecting the posterior corners of the eyes, may be present; on flanks there may be several oval dark spots, more or less arranged in two longitudinal rows. Females with a light and dark brown pattern, with radiating dark lines from the eyes, the dorsal ones meeting each other middorsally, while lateral ones may extend along the neck; antehumeral bar less conspicuous than in males; a vertebral light band, with irregular borders, is more or less evident; oval dark spots may be present on flanks. Tail, in males and females, with dark brown transverse bands dorsally; female RMNH 26417 with rather distinct dark and light brown rings distally on tail. Regenerated tail mostly uniformly brown. Juvenile pattern similar to that of females (Avila-Pires 1995: 273 ff). 
CommentSynonymy partly after KLUGE 1993. Donoso-Barros, 1968 synonymized G. ferrugineus with G. humeralis, without justification. Pinto et al. 2018 resurrected the name G. ferrugineus for the Trinidad population, which is genetically clearly distinguishable from mainland G. humeralis. However, G. ferrugineus is closely related to mainland G. humeralis and “Gonatodes ferrugineus is currently morphologically indistinguishable from G. humeralis although there appear to be some qualitative differences in proportionality of the face, body size, and coloration in adult males that may, upon further investigation, diagnose this species”. Hence we leave it within G. humeralis for the time being.
“Adult [G. ferrugineus] males from Trinidad are generally not as colorful as those from mainland South America (Supplemental Fig. 1). Trinidadian males lack red spots on the sides of the body and their heads tend to favor orange/yellow rather than red and white/blue, both of which are typical features of most South American populations” (Pinto et al. 2018).

Synonymy (Cnemaspis timoriensis): Cnemaspis timoriensis was reported from Indonesia (Timor), a locality that must be in error, given that C. timoriensis is a synonym of G. humeralis (Rösler et al. 2019). Cnemaspis timoriensis has priority over G. humeralis, but Rösler et al. 2019: 501 suggested to apply to the ICZN under Article 23.9.3 for suppression of Cnemaspis timoriensis. Cnemaspis timoriensis is only known from 2 specimens (Rösler 2016), namely the alleged type and a specimen in MTD, which is actually not G. humeralis (Rösler et al. 2019). Bauer 2013 stated that the specimens are “probably not a member of this genus” (Cnemaspis). Rösler 2016 concluded that the MNHN specimen belongs to Garthia or Homonota and the ZMB specimen is a Gonatodes. A.M. Bauer (pers. comm., 22 July 2017) suggested to delete this “species”. Ineich (in press) synonymized this species with XYZ (pers. comm., March 2019).

Distribution: not on Tobago (Murphy 1997), although reported from there by MERTENS 1972. Populations on Trinidad represent G. ferrugineus, a closely related sister species of G. humeralis. However, because both are closely related, epsecially given the relationships among other Gonatodes (Fig. 2 in Pinto et al., despite a 9.4% pairwise divergence at the mitochondrial locus ND2) and morphologically indistinguishable (Pinto et al. 2018) we leave ferrugineus within G. humeralis. G. ferrugineus may occur in Venezuela too, but Pinto et al. 2018 have not sampled this area.

Behavior: mainly diurnal

Sympatry: Gonatodes humeralis occurs in sympatry with many larger species of the genus, e.g. G. annularis, G. concinnatus, G. hasemani, G. tapajonicus (Avila-Pires 1995; Dixon & Soini 1975; Rivero-Blanco 1979; Moravec et al. 2001), and G. nascimentoi. 
References
  • Aguiar, Kurazo M. Okada and Carlos E. Costa-Campos. 2014. Gonatodes humeralis (Trinidad gecko) predation. Herpetological Review 45 (3): 498-499 - get paper here
  • Anaissi,Costa- Jessica Stefany 2021. Record of tail bifurcation and hindlimb malformation in Gonatodes humeralis (Guichenot, 1855) (Squamata, Gekkonidae) from the Amazonian region of northern Brazil. Herpetology Notes 14: 341-343 - get paper here
  • Araújo, Jailini da Silva; Vinicius Guerra, Moises Barbosa de Souza, Fabiano Corrêa 2020. Assemblage of lizards of a forest fragment in southwest Amazon, Brazil. Herpetology Notes 13: 895-901 - get paper here
  • Ascenso, A.C., De Oliveira, L.S. & Filho, F.C. 2017. Gonatodes humeralis (Trinidad Gecko) Endoparasitism. Herpetological Review 48 (3): 644-645. - get paper here
  • Auguste, Renoir J. 2019. Herpetofaunal checklist for six pilot protected areas in Trinidad and Tobago. Herpetology Notes 12: 577-585 - get paper here
  • Avila-Pires, T.C.S. 1995. Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen 299: 1-706 - get paper here
  • Avila-Pires, Teresa C. S.; Daniel G. Mulcahy, Fernanda P. Werneck, and Jack W Sites 2012. Phylogeography of the Teiid Lizard Kentropyx calcarata and the Sphaerodactylid Gonatodes humeralis (Reptilia: Squamata): Testing A Geological Scenario for the Lower Amazon–Tocantins Basins, Amazonia, Brazil. Herpetologica 68 (2): 272-287. - get paper here
  • Avila-Pires, Teresa C.S.; Kleiton R. Alves-Silva, Laís Barbosa, Fabrício S. Correa, Jorge F. A. Cosenza, Ana Paula V. Costa-Rodrigues, Aurea A. Cronemberger, Marinus S. Hoogmoed, Geraldo R. Lima-Filho, Adriano O. Maciel, Alexandre F. R. Missassi, Lyw 2018. Changes in Amphibian and Reptile Diversity Over Time in Parque Estadual do Utinga, a Protected Area Surrounded by Urbanization. Herpetology Notes 11: 499-512 - get paper here
  • Barrio-Amorós, C.L. 2010. GONATODES - Los Geckos diurnos de Venezuela. RioVerde 2010; 43-50
  • BARRIO-AMORÓS, CESAR L. & JUAN C. ORTÍZ 2016. Venezuelan geckos (Gekkonidae, Phyllodactylidae, Sphaerodactylidae) in the collection of the Universidad de Concepción in Chile, with description of the type series of Gonatodes ligiae and Gonatodes petersi (Sphaerodactylidae). Zootaxa 4136 (3): 537–552 - get paper here
  • Bauer, A.M.; Günther,R. & Klipfel,M. 1995. The herpetological contributions of Wilhelm C.H. Peters (1815-1883). SSAR Facsimile Reprints in Herpetology, 714 pp.
  • Bernarde, Paulo Sérgio;, Reginaldo Assêncio Machado & Luiz Carlos Batista Turci 2011. Herpetofauna da área do Igarapé Esperança na Reserva Extrativista Riozinho da Liberdade, Acre – Brasil. Biota Neotropica 11 (3): 117-144 - get paper here
  • Boos, H.E.A. 1984. The terrestrial reptiles of Monos island. Living World - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Bour, Roger & Édouard-Raoul Brygoo 2013. Séraphin Braconnier (1812-1884), le premier «garçon de laboratoire» de la chaire des Reptiles et Poissons du Muséum de Paris. Bull. Soc. Herp. France 148: 503-513
  • Burt, Charles E. & Myers, George S. 1942. Neotropical lizards in the collection of the Natural History Museum of Stanford University. Stanford University Press, 52 pp.
  • CATENAZZI, A., LEHR, E. & VON MAY, R. 2013. The amphibians and reptiles of Manu National Park and its buffer zone, Amazon basin and eastern slopes of the Andes, Peru. Biota Neotropica 13 (4): 269-283
  • Cole, C.J. & Kok, P.J. 2006. New Species of Gekkonid Lizard (Sphaerodactylinae: Gonatodes) from Guyana, South America. American Museum Novitates (3524): 1-16 - get paper here
  • Cole, Charles J.; Carol R. Townsend, Robert P. Reynolds, Ross D. MacCulloch, and Amy Lathrop 2013. Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis. Proceedings of the Biological Society of Washington 125 (4): 317-578; plates: 580-620 - get paper here
  • Cooper Jr., W.E. 2005. Duration of movement as a lizard foraging movement variable. Herpetologica 61 (4): 363-372 - get paper here
  • Cope, E.D. 1863. Descriptions of new American Squamata in the Museum of the Smtihsonian Institution. Proc. Acad. Nat. Sci. Philadelphia 15 [1863]: 100-106 - get paper here
  • Das I 1994. Cnemaspis gordongekkoi, a new gecko from Lombok, Indonesia, and the biogeography of oriental species of Cnemaspis (Squamata: Sauria: Gekkonidae). Hamadryad 18: 1-9 [1993] - get paper here
  • Das, I. 2005. Revision of the Genus Cnemaspis Strauch, 1887 (Sauria: Gekkonidae), from the Mentawai and Adjacent Archipelagos off Western Sumatra, Indonesia, with the Description of Four New Species. Journal of Herpetology 39 (2): 233–247 - get paper here
  • Das, I. & Grismer, L.L. 2003. Two new species of Cnemaspis Strauch 1887 (Squamata, Gekkonidae) from the Seribuat Archipelago, Pahang and Johor States, West Malaysia. Herpetologica 59 (4): 544-552 - get paper here
  • de Figueiredo, Vinícius A. M. B.; Souza, Naziel S.; Anaissi, Jessica S. C.; Sanches, Patrick R.; Costa-Campos, Carlos E. 2020. Diet of four lizards from an urban forest in an area of Amazonian biome, Eastern Amazon. Cuad. herp. 34 (2): - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Diele-Viegas, L. M., Werneck, F. P. & Rocha, C. F. D. 2019. Climate change effects on population dynamics of three species of Amazonian lizards. Comp. Biochem. Physiol. Part A Mol. Integr. Physiol. 236, 110530 - get paper here
  • Dixon, J. R.; Soini, P. 1975. The reptiles of the upper Amazon basin, Iquitos region, Peru, Part I. Lizards and Amphisbaenians. Milwaukee Public Museum Contributions in Biology and Geology 4: 1-58 [1986?] - get paper here
  • Duellman, W. E. 2005. Cusco Amazónico: The Lives of Amphibians and Reptiles in an Amazonian Rainforest. Comstock Pub Assoc.
  • Duellman, W.E., & Salas, A.W. 1991. Annotated checklist of the amphibians and reptiles of Cuzco Amazonico, Peru. Occas. Papers Mus. of Natur. Hist., Univ. of Kansas, Lawrence (143): 13 pp. - get paper here
  • Duméril, A.M. C. and G. Bibron. 1836. Erpetologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 3. Libr. Encyclopédique Roret, Paris, 528 pp. - get paper here
  • Ferreira, A. S. 2021. Rapid survey of the herpetofauna of Estação Ecológica Alto Maués: a rarely accessed area in the Brazilian Amazonia. Biota Amazônia 11(1): 22-28 - get paper here
  • Fonseca, Wirven Lima da; Julivaldo Dantas da Silva, Arthur Diesel Abegg, Conrado Mario da Rosa, Paulo Sergio Bernarde 2019. Herpetofauna of Porto Walter and surrounding areas, Southwest Amazonia, Brazil. Herpetology Notes 12: 91-107 - get paper here
  • Freitas, M. A. de, Venancio, N. M., Abegg, A. D., Azevedo, W. S, Pereira, V. O., Zanotti, A. P., Veloso, A., Schwarzbach, L., Oliveira, A. G., da Silva, R. C. C., de Amorim, V. R. G. and de Moura, G. J. B. 2020. Herpetofauna at the Rio Acre Ecological Station, Amazon Rainforest, Brazil. Herpetology Notes 13: 33-48. - get paper here
  • Fugler,C. M. 1983. Lista preliminar de los anfibios y reptiles de Tumi Chucua. Mus. Nac. Hist. Nat. (Bolivia) Comunicación 2: 4-11.
  • Gamble, Tony; Andrew M. Simons, Guarino R. Colli and Laurie J. Vitt 2008. Tertiary climate change and the diversification of the Amazonian gecko genus Gonatodes (Sphaerodactylidae, Squamata). Molecular Phylogenetics and Evolution 46 (1): 269-277 - get paper here
  • Gardner, T.A., Ribeiro Junior, M.A., BarlowJ., Avila-Pires, T.C.S., Hoogmoed, M.S. & Peres, C. 2007. The Value of Primary, Secondary, and Plantation Forests for a Neotropical Herpetofauna. Conservation Biology 21 (3): 775–787 - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here
  • Gorzula, Stefan & Senaris, J. Celsa 1999. In: Contribution to the herpetofauna of the Venezuelan Guayana. I: a data base. Scientia Guaianae, Caracas, No. 8 [1998], 269+ pp.; ISBN 980-6020-48-0
  • Grismer, L.L. & Das, I. 2006. A new species of gekkonid lizard of the genus Cnemaspis STRAUCH 1887 from Pulau Pemanggil, Johor, West Malaysia. Herpetological Natural History 10 (1): 1-7 - get paper here
  • Guichenot, A. 1855. Animaux nouveaux ou rares recueillis pendant l’Expédition dans les parties centrales de l’Amérique du Sud, de Rio de Janeiro a Lima, et de Lima au Para; Exécutée par ordre du Gouvernement Francais pendant les années 1843 a 1847, sous la direction du P. Bertrand, Paris. - get paper here
  • Harvey, M. B., APARICIO, J., CORTEZ, C., GONZÁLES, L., GUERRA, J. F., MONTAÑO, M. E. & PÉREZ, M. E. 1998. Reptile and amphibian species of Parque Nacional Noel Kempff Mercado. In: Killeen, T. J. & T. S. Schulenberg (eds). A biological assessment of Parque Nacional Noel Kempff Mercado, Bolivia. Conservation International Washington. Rap Working Papers 10: 348-355.
  • Hedges SB, Powell R, Henderson RW, Hanson S, and Murphy JC 2019. Definition of the Caribbean Islands biogeographic region, with checklist and recommendations for standardized common names of amphibians and reptiles. Caribbean Herpetology 67: 1–53
  • Hluschi, D. 2009. Gonatodes - die unbekannten Juwelen der Neotropis. Reptilia (Münster) 14 (78): 16-23 - get paper here
  • Hoogmoed M.S.; de Avila-Pires T C S 1991. Annotoated checklist of the herpetofauna of Petit Saut, Sinnamary River, French Guiana. Zoologische Mededelingen 65: 53-88 - get paper here
  • Hoogmoed, M.S., & Lescure, J. 1975. An annotated checklist of the lizards of French Guiana, mainly based on two recent collections. Zoologische Mededelingen 49(13): 141-172. - get paper here
  • Johnson, M.L. 1946. Herpetological Notes from Trinidad. Copeia 1946 (2): 108 - get paper here
  • Kawashita-Ribeiro, Ricardo Alexandre; Jacqueline Pimentel Silva, Anelise Figueiredo da Silva, Luana Aparecida Gomes de Arruda, Tamí Mott, Marcos André de Carvalho 2011. Descobrindo a Amazônia Meridional: Biodiversidade da Fazenda São Nicolau Os Répteis Escamosos (Reptilia, Squamata)da Fazenda São Nicolau, Cotriguaçu,Mato Grosso, Brasil, um Estudo Preliminar. In: Descobrindo a Amazônia Meridional: biodiversidade da F Paue Prosa Comunicação Ltda. - get paper here
  • Kornacker, Paul M.;Meyer zu Heyde, Christian 1998. Herpetologische Eindrücke einer Venezuelareise Teil 3: El Embalse de Guri - der Guri-Stausee. Elaphe 6 (4): 61-69
  • Lehr, E. 2002. Amphibien und Reptilien in Peru. Natur und Tier-Verlag (Münster), 208 pp. - get paper here
  • Lima Moraes, L. J. C. de, do Nascimento Rainha, R., de Pinho Werneck, F., de Souza Oliveira, A. F., Gascon, C., & de Carvalho, V. T. 2022. Amphibians and reptiles from a protected area in western Brazilian Amazonia (Reserva Extrativista do Baixo Juruá). Papéis Avulsos de Zoologia, 62, e202262054-e202262054 - get paper here
  • LiVigni, F. (ed.) 2013. A Life for Reptiles and Amphibians, Volume 1. Chimaira, Frankfurt, 495 pp. - get paper here
  • Madella-Auricchio, Cláudia Renata; Paulo Auricchio Enio Saraiva Soares 2017. Reptile species composition in the Middle Gurguéia and comparison with inventories in the eastern Parnaíba River Basin, State of Piauí, Brazil. Papéis Avulsos de Zoologia 57(28): 375-386, - get paper here
  • Mendes-Pinto, T. J. & S. Marques de Souza 2011. Preliminary assessment of amphibians and reptiles from Floresta Nacional do Trairão, with a new snake record for the Pará state, Brazilian Amazon. Salamandra 47 (4): 199-206 - get paper here
  • Mertens, R. 1972. Herpetofauna tobagana. Stuttgarter Beitr. zur Naturkunde nr. 252 22 pp. - get paper here
  • Metcalf, Matthew; Alexander Marsh, Emerson Torres, Devon Graham, Charles Gunnels 2020. Herpetofauna of the Santa Cruz Forest Preserve in the Peruvian Amazon Basin. Herpetology Notes 13: 753-767 - get paper here
  • Miranda, Daniele Bazzo; Nathocley Mendes Venâncio, Saymon de Albuquerque 2014. Rapid survey of the herpetofauna in an area of forest management in eastern Acre, Brazil. Check List 10 (4): 893-899 - get paper here
  • Miranda, J.P. & G.V. Andrade 2003. Seasonality in Diet, Perch Use, and Reproduction of the Gecko Gonatodes humeralis from Eastern Brazilian Amazon Journal of Herpetology 37 (2): 433-438. - get paper here
  • Morato, S.A.A.; Calixto, P.O.; Mendes, L.R.; Gomes, R.; Galatti, U.; Trein, F.L.; Oliveira, F.S.; Ferreira, G.N. 2014. Guia fotográfico de identificação da herpetofauna da Floresta Nacional de Saracá-Taquera, Estado do Pará. Curitiba: STCP Engenharia de Projetos Ltda.; Porto Trombetas: MRN – Mineração Rio do Norte S.A.; 213 p. - get paper here
  • Morato, Sérgio Augusto Abrahão; Guilherme Nunes Ferreira; Michela Rossane Cavilha Scupino (eds.) 2018. Herpetofauna da Amazônia Central: Estudos na FLONA de Saracá-Taquera. Curitiba, Pr: STCP Engenharia de Projetos Ltda.; Porto Trombetas, Pa: MRN – Mineração Rio do Norte S.A., 2018.<br />210p. - get paper here
  • Moravec J., Aparicio J. 2005. Notes on the herpetofauna of Bioceanica and Bolpebra (Provincia Nicolas Suaréz, Departamento Pando, Biolivia). Journal of the National Museum (Prague), Natural History Series 174 (1-4): 95-113
  • Oliveira, D.P.; S.M. Souza; L. Frazão; A.P. Almeida; T. Hrbek 2014. Lizards from central Jatapú River, Amazonas, Brazil. Check List 10 (1): 46-53 - get paper here
  • Oliveira, F. R. C. de, Passos, D. C., & Borges-Nojosa, D. M. 2021. Ecology of the lizard Gonatodes humeralis (Sphaerodactylidae) in a coastal area of the Brazilian semiarid: What differs from the Amazonian populations?. Journal of Arid Environments, 190, 104506 - get paper here
  • O’Shaughnessy,A.W.E. 1876. Descriptions of new species of Gekkotidae in the British Museum Collection. Ann. Mag. Nat. Hist. (4) 16: 262-266 - get paper here
  • Pedroso-Santos, Fillipe; Patrick Ribeiro Sanches, and Carlos Eduardo Costa-Campos, 2019. Anurans and reptiles of the Reserva Extrativista Beija-Flor Brilho de Fogo, Amapá state, eastern Amazon. Herpetology Notes 12: 799-807 - get paper here
  • Peters, Wilhem Carl Hartwig 1871. Über eine von Hrn. Dr. Robert Abendroth in dem Hochlande von Peru gemachte Sammlung von Amphibien, welche derselbe dem Königl. zoologischen Museum geschenkt hat. Monatsber. königl. Akad. Wiss. Berlin. 1871 (August): 397-404 - get paper here
  • Pianka, E.R. & Vitt, L.J. 2003. Lizards - Windows to the Evolution of Diversity. University of California Press, Berkeley, 347 pp. [review in Copeia 2004: 955] - get paper here
  • Pinto, B. J., Colli, G. R., Higham, T. E., Russell, A. P., Scantlebury, D. P., Vitt, L. J., & Gamble, T. 2018. Population genetic structure and species delimitation of a widespread, Neotropical dwarf gecko. Molecular Phylogenetics and Evolution 133: 54-66 [2019] - get paper here
  • Pogoda, P. 2018. Auf Expedition in der „verlorenen Welt“ – herpetologische Eindrücke aus dem Naturreservat Iwokrama in Zentral-Guyana. Reptilia 23 (129): 70-79
  • Prudente, A.L.C.; F. Magalhães; A. Menks; J.F.M. Sarmento. 2013. Checklist of Lizards of the Juruti, state of Pará, Brazil. Check List 9 (1):42-50 - get paper here
  • Rabosky, Daniel L.; Rudolf von May, Michael C. Grundler and Alison R. Davis Rabosky 2019. The Western Amazonian Richness Gradient for Squamate Reptiles: Are There Really Fewer Snakes and Lizards in Southwestern Amazonian Lowlands? Diversity 11: 199; doi:10.3390/d11100199 - get paper here
  • RIBEIRO-JÚNIOR, MARCO A. 2015. Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. II. Gekkonidae, Phyllodactylidae, Sphaerodactylidae. Zootaxa 3981 (1): 001–055 - get paper here
  • Ribeiro-Júnior, Marco A. & Silvana Amaral 2016. Diversity, distribution, and conservation of lizards (Reptilia: Squamata) in the Brazilian Amazonia. Neotropical Biodiversity, 2:1, 195-421 - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Roberto, Igor Joventino; Thieres Pinto, Álcio Schlickmann and Andressa Fraga. 2014. From Amazonia to the semi-arid: the unexpected record of Gonatodes humeralis (Squamata: Sphaerodactylidae) for the Caatinga Biome. Herpetology Notes 7: 309-311 - get paper here
  • Rodrigues-França, Frederico Gustavo & Nathocley Mendes-Venâncio 2010. Reptiles and amphibians of a poorly known region in southwest Amazonia Biotemas, 23 (3): 71-84 - get paper here
  • Rösler H 2016. Bemerkungen über einige Geckoarten der Zoologischen Staatssammlung München. Gekkota Suppl. 3: 1-94
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, Herbert 2017. Gecko-Chorologie (Squamata: Gekkota). Gekkota (4): 1-160
  • RÖSLER, HERBERT; L. LEE GRISMER, IVAN INEICH & HINRICH KAISER, 2019. The purportedly Indo-Australian gecko species Cnemaspis timoriensis (Duméril & Bibron, 1836) is actually the first named species of the neotropical genus Gonatodes Fitzinger, 1843 (Squamata: Gekkonidae). Zootaxa 4576 (3): 483–509 - get paper here
  • Russell, A.P., Garner, A.M. 2023. Solutions to a Sticky Problem: Convergence of the Adhesive Systems of Geckos and Anoles (Reptilia: Squamata). In: Bels, V.L., Russell, A.P. (eds) Convergent Evolution. Fascinating Life Sciences. Springer, Cham - get paper here
  • Santos, Marina Meireles dos; Robson Waldemar Ávila and Ricardo Alexandre Kawashita-Ribeiro. 2011. Checklist of the amphibians and reptiles in Nobres municipality, Mato Grosso state, central Brazil. Herpetology Notes 4: 55-461 - get paper here
  • Schmidt, K. E, & INGER, R. F. 1951. Amphibians and reptiles of the Hopkins-Branner expedition to Brazil. Fieldiana: Zoology 31: 439-465 - get paper here
  • Señaris, J. Celsa; María Matilde Aristeguieta Padrón, Haidy Rojas Gil y Fernando J. M. Rojas-Runjaic 2018. Guía ilustrada de los anfibios y reptiles del valle de Caracas, Venezuela. Ediciones IVIC, Instituto Venezolano de Investigaciones Científicas (IVIC). Caracas, Venezuela. 348 pp.
  • Silva, F.M.; A.C. Menks; A.L.C. Prudente; J.C.L. Costa; A.E.M. Travassos; U. Galatti. 2011. Squamate Reptiles from municipality of Barcarena and surroundings, state of Pará, north of Brazil. Check List 7 (3): 220-226 - get paper here
  • Silva, Marcélia Basto da; Wáldima Alves da Rocha, Janete Diane Nogueira-Paranhos 2016. Checklist of reptiles of the Amazonia-Caatinga-Cerrado ecotonal zone in eastern Maranhão, Brazil. Herpetology Notes 9: 7-14 - get paper here
  • STURARO, MARCELO JOSÉ & TERESA C.S. AVILA-PIRES 2011. Taxonomic revision of the geckos of the Gonatodes concinnatus complex (Squamata: Sphaerodactylidae), with description of two new species. Zootaxa 2869: 1–36 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Uchôa LR, Delfim FR, Mesquita DO, Colli GR, Garda AA, Guedes TB 2022. Lizards (Reptilia: Squamata) from the Caatinga, northeastern Brazil: Detailed and updated overview. Vertebrate Zoology 72: 599-659 - get paper here
  • Ulmer, P. 2007. EIn Aufenthalt in einem abgelegenen Teil des brasilianischen Amazonasgebiets: herpetologisches Tagebuch. Reptilia (Münster) 12 (66): 42-50 - get paper here
  • Vaz-Silva, W.; RM Oliveira, AFN Gonzaga, KC Pinto, FC Poli, TM Bilce, M Penhacek, L Wronski, JX Martins, TG Junqueira, LCC, Cesca VY, Guimarães RD. Pinheiro 2015. Contributions to the knowledge of amphibians and reptiles from Volta Grande do Xingu, northern Brazil Braz. J. Biol., 75 (3) (suppl.): S205-S218 - get paper here
  • Vitt, L. J.; Zani, P. A.;Barros, A. Monteiro de 1997. Ecological variation among populations of the gekkonid lizard Gonatodes humeralis in the Amazon Basin. Copeia 1997 (1): 32-43 - get paper here
  • Vitt, L.; Magnusson, W.E.; Ávila-Pires, T.C. & Lima, A.P. 2008. Guide to the Lizards of Reserva Adolpho Ducke, Central Amazonia. Attema, Manaus, 180 pp.
  • Waldez, F., Menin, M., & Vogt, R. C. 2013. Diversidade de anfíbios e répteis Squamata na região do baixo rio Purus, Amazônia Central, Brasil. Biota Neotropica, 13: 300–316 - get paper here
  • Wermuth, H. 1965. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich (80): 1—246
  • Whithworth, A. & Beirne, C. 2011. Reptiles of the Yachana Reserve. Global Vision International, 130 pp. - get paper here
 
External links  
Is it interesting? Share with others:

As link to this species use URL address:

https://reptile-database.reptarium.cz/species?genus=Gonatodes&species=humeralis

without field 'search_param'. Field 'search_param' is used for browsing search result.



Please submit feedback about this entry to the curator