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Gonatodes timidus KOK, 2011

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Higher TaxaSphaerodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common Names 
SynonymGonatodes timidus KOK 2011 
DistributionC Guyana (Iwokrama Forest Reserve)

Type locality: base of Iwokrama Mountains, Potaro-Siparuni District, Guyana (04° 19’ 52”N, 058° 47’ 58”W, 209 m elevation).  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: IRSNB 2664 
DiagnosisDiagnosis: The new taxon is mainly distinguished from all known congeners by the following combination of characters: moderate body size, absence of a clearly differentiated elongate supraciliary spine, absence of clusters of distinctly enlarged conical scales on sides, escutcheon scales on posterior belly and ventral surfaces of thighs in males evident, three to four lateral rows of scales on distal part of fingers and toes, medial subcaudal scales distinctly differentiated from adjacent scales on non-regenerated tail (in a 1’1” sequence), and obvious sexual dichroma- tism, the males having the upper surface of the head black with bluish white to vivid yellow irregular stripes and blotches. 
CommentBehavior: diurnal

Sympatry: sympatric, but never syntopic, with Gonatodes alexandermendesi.

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific name is considered a noun in apposition and is derived from the Latin adjective timidus meaning “shy” or “fearful”. It refers to the tendency of individuals of the new species to avoid to be seen by hastily escaping between rocks, making them very difficult to collect. 
  • Cole, Charles J.; Carol R. Townsend, Robert P. Reynolds, Ross D. MacCulloch, and Amy Lathrop 2013. Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis. Proceedings of the Biological Society of Washington 125 (4): 317-578; plates: 580-620 - get paper here
  • KOK, PHILIPPE J. R. 2011. A new species of the genus Gonatodes Fitzinger, 1843 (Reptilia: Sphaerodactylidae) from central Guyana, northern South America. Zootaxa 3018: 1–12 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
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