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Goniurosaurus varius QI, GRISMER, LYU, ZHANG, LI & WANG, 2020

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Higher TaxaEublepharidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Nanling Leopard Gecko]
Chinese: 南岭睑虎 
SynonymGoniurosaurus varius QI, GRISMER, LYU, ZHANG, LI & WANG 2020 
DistributionChina (N Guangdong: Nanling National Nature Reserve)

Type locality: Nanling National Nature Reserve (ca 560 m a.s.l.), Chengjia Yao Ethnic Township, Yangshan County, Guangdong Province, China.  
Reproduction 
TypesHolotype. SYS r002333, adult male (Fig. 7), collected by Liang Zhang on 20 September 2019. Paratypes. One adult male (SYS r002331) and three adult females (SYS r002330, SYS r002362–2363), collected by Zhi-Ren Zhang, Yu Zhang, and Peng Cen on 6 August 2018, from Nanling National Nature Reserve, Chengjia Yao Ethnic Township at eleva- tions between 180 and 560 m. 
DiagnosisDiagnosis: Goniurosaurus varius sp. nov. can be distinguished from all other congeners by the combination of the following characters: (1) adult body size moderate, measuring 81.5–86.3 mm in SVL; (2) nasal scales surrounding nares 7–9; (4) internasal usually single, rarely two; (5) eyelid fringe scales 50–56; (6) granular scales of the upper eyelids similar in size to those on the top of the head; (7) scales around midbody 101–110; (8) dorsal tubercle rows at midbody 21–24; (9) paravertebral tubercles between limb insertions 27–29; (10) claws sheathed by four scales, dorsal scale small, two lateral scales short and shell-shaped; (11) axillary pockets deep; (12) presence of ten precloacal pores in males, and absent in females; (13) dorsal ground color of head, body, and limbs in adults reddish brown, mottled with varied spots and stripes; (14) nuchal loop usually incomplete, if complete, posteriorly rounded; (15) presence of four thin dorsal body bands with dark spots, bordered with black anteriorly and posteriorly, sometime last two bands indistinct; (16) usually presence of a longitudinal light colored vertebral stripe on the trunk of body; (17) light pink beneath, with dark brown lateral spots; (18) iris orange-red.


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CommentDistribution: see map in Ngo et al. 2023. 
EtymologyThe specific name varius means varied or diverse in Latin and refers to its variable dorsal color pattern. 
References
  • Ngo, H. N., Rödder, D., Grismer, L., Nguyen, T. Q., Le, M. D., Qi, S., & Ziegler, T. 2023. Extraordinary diversity among allopatric species in the genus Goniurosaurus (Squamata: Eublepharidae): understanding niche evolution and the need of conservation measures. Biodiversity and Conservation, 1-23 - get paper here
  • Qi S, Grismer LL, Lyu Z-T, Zhang L, Li P-P, Wang Y-Y 2020. A definition of the Goniurosaurus yingdeensis group (Squamata, Eublepharidae) with the description of a new species. ZooKeys 986: 127-155 - get paper here
  • Zhou, Z., Ding, L., Liu, Z., Fu, L., Xu, L., Feng, L., ... & Zhou, Y. 2023. The complete mitochondrial genome of Goniurosaurus varius (Squamata: Eublepharidae). Mitochondrial DNA Part B, 8(11), 1215-1219 - get paper here
 
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