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Goniurosaurus hainanensis BARBOUR, 1908

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Higher TaxaEublepharidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesG: Hainan-Krallengecko
Chinese: 海南睑虎 
SynonymGoniurosaurus hainanensis BARBOUR 1908: 316
Goniurosaurus hainanensis — GRISMER 2000
Goniurosaurus hainanensis — PIANKA & VITT 2003: 171
Goniurosaurus hainanensis — DICKHOFF 2004
Goniurosaurus hainanensis — SEUFER et al. 2005
Goniurosaurus lichtenfelderi hainanensis 
DistributionSE-Hainan (China)

Type locality: Mt. Wuchi, Central Hainan.  
TypesHolotype: MCZ 7104 
DiagnosisOriginal diagnosis (genus): “Digits moderate; otherwise exactly as in Aelurosaurus. Body covered with excessively small, flat juxtaposed scales and larger tubercle-like scales. Upper and lower eyelids well developed, as in Aelurosaurus (Geckonidae) and in the Eublepharidae. Pupil vertical. Tail elongate with whorls of scales proximally. That the tail is capable of being curled up is evident from its position in the type preserved in alcohol. Possibly this genus should also contain Pentadactylas brunneus Cope, which Boulenger placed provisionally in Aelurosaurus.” (from BARBOUR 1908).

Diagnosis (genus): rod-like clavicles; the anterior portion of interclavicle being dorsal to clavicles; a cleft rostral scale; enlarged caudal scale whorls; the anterolateral palatine projections being overlapped by the maxillae; the posteroventral portion of the parietal being smooth; the posterior section of the prootic wings not contacting the medial bulge of the prootics; the lack of tubercles surrounding the auditory meatus; extensive distal fusion of sacral transverse processes; a dorsal ridge on the ulnare; and the retention of the juvenile caudal color pattern of black and white bands into adulthood, which are unique to Goniurosaurus (after Grismer 1988, and Grismer et al. 1999, cited in WANG et al. 2010). 
CommentSynonymy: Goniurosaurus hainanensis has been synonymized with G. lichtenfelderi by SMITH 1935.

Type Species: Goniurosaurus hainanensis BARBOUR 1908 is the type species of the genus Goniurosaurus BARBOUR 1908.

Phylogenetics: for a phylogenetic analysis of the genus see Liang et al. 2018. 
  • Barbour, THOMAS 1908. Some new reptiles and amphibians. Bull. Mus. Comp. Zool. Harvard 51 (12): 315-325 - get paper here
  • Barbour, Thomas 1909. Notes on Amphibia and Reptilia from Eastern Asia. Proc. New England zool. Club 4: 53-78, 2 plates - get paper here
  • Blair, Christopher; Nikolai L. Orlov, Hai-tao Shi, and Robert W. Murphy 2009. A Taxonomic Re-Evaluation of Goniurosaurus hainanensis (Squamata: Eublepharidae) from Hainan Island, China. Russ. J. Herpetol. 16 (1): 35-40 - get paper here
  • Chen, Tian-Bo; Yuan-Jun Meng, Ke Jiang,, Pi-Peng Li, Bo-Hao Wen, Wenhua Lu, James Lazell, and Mian Hou 2014. New Record of the Leopard Gecko Goniurosaurus araneus (Squamata: Eublepharidae) for China and habitat partitioning between geographically and Phylogenetically Close Leopard Geckos. IRCF Reptiles & Amphibians 21 (1): 16–27
  • Dickhoff, A. 2004. Ein Tiger im Terrarium - Haltung und Nachzucht von Goniurosaurus araneus (GRISMER, VIETS & BOYLE 1999). Draco 5 (18): 76-81 - get paper here
  • Grismer, L. Lee 2002. Goniurosaurus: Ancient Gekkos of the Far East. Gekko 3 (1): 22-28
  • Grismer, L. Lee, Brian E. Viets and Lawrence J. Boyle. 1999. Two new continental species of Goniurosaurus (Squamata: Eublepharidae) with a phylogeny and evolutionary classification of the genus. Journal of Herpetology 33 (3): 382-393 - get paper here
  • Grismer, L.L. 1988. Phylogeny, taxonomy, classification, and biogeography of eublepharid geckos. in Estes, R., & Pregill, G., eds., Phylogenetic relationships of the lizard families: essays commemorating Charles L. Camp (Stanford Univ. Press, Stanford): 369-469.
  • Grismer,L.L. 2000. Goniurosaurus murphyi ORLOV & DAREVSKY: a junior synonym of Goniurosaurus lichtenfelderi Mocquard. Journal of Herpetology 34 (3): 486-488 - get paper here
  • Johann, H. 2006. Pfelge und Nachzucht von Goniurosaurus hainanensis BARBOUR 1908. Elaphe 14 (4): 27-31
  • Liang, B., Zhou, R.-B., Liu, Y.-L., Chen, B., Lee Grismer, L., & Wang, N. 2018. Renewed classification within Goniurosaurus (Squamata: Eublepharidae) uncovers the dual roles of a continental island (Hainan) in species evolution. Molecular Phylogenetics and Evolution - get paper here
  • Pianka, E.R. & Vitt, L.J. 2003. Lizards - Windows to the Evolution of Diversity. University of California Press, Berkeley, 347 pp. [review in Copeia 2004: 955] - get paper here
  • Seufer, H.; Y. Kaverkin & A. Kirschner (eds.) 2005. Die Lidgeckos. Kirschner und Seufer Verlag, 238 pp.
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Wang, Ying-Yong; Jian-Huan Yang, and L. Lee Grismer 2013. A New Species of Goniurosaurus (Squamata: Eublepharidae) from Libo, Guizhou Province, China. Herpetologica 69 (2): 214-226. - get paper here
  • Wang, Ying-Yong; Jian-Huan Yang, and Rong-Feng Cui 2010. A NEW SPECIES OF GONIUROSAURUS (SQUAMATA: EUBLEPHARIDAE) FROM YINGDE, GUANGDONG PROVINCE, CHINA. Herpetologica 66 (2): 229 - get paper here
  • Wang, Ying-Yong; Meng-Jie Jin, Yu-Long Li, and L. Lee Grismer 2014. Description of a New Species of Goniurosaurus (Squamata: Eublepharidae) from the Guangdong Province, China, Based on Molecular and Morphological Data. Herpetologica Sep 2014, Vol. 70, No. 3: 309-322. - get paper here
  • ZHOU, RUN-BANG; NING WANG, BEI CHEN, BIN LIANG 2018. Morphological evidence uncovers a new species of Goniurosaurus (Squamata: Eublepharidae) from the Hainan Island, China. Zootaxa 4369 (2): 281–291 - get paper here
  • ZHU, XIAO-YU; CHU-ZE SHEN, YUN-FEI LIU, LIN CHEN, ZHENG LI, & ZHU-QING HE 2020. A new species of Goniurosaurus from Hainan Island, China based on molecular and morphological data (Squamata: Sauria: Eublepharidae). Zootaxa 4772 (2): 349–360 - get paper here
  • Ziegler, T.; Truong, N.Q.; Schmitz, A.; Stenke, R. & Rösler, H. 2008. A new species of Goniurosaurus from Cat Ba Island, Hai Phong, northern Vietnam (Squamata: Eublepharidae). Zootaxa 1771: 16–30 - get paper here
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